Physaria newberryi
Physaria didymocarpa
Newberry twinpod, Newberry's twinpod
common twinpod, double twinpod, twin bladderpod
several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm.
several from base, decumbent, (unbranched, leafy for the genus), ca. 1 dm.
(ascending to erect, petiole slender);
blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse).
(forming a strong rosette; long-petiolate);
blade obovate, 1.5–4(–8) cm, (base ± abruptly narrowing to petiole), margins usually repand or dentate, rarely entire, (apex usually angular, surfaces silvery).
blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire.
blade oblanceolate, 1–2 cm (width 4–8 mm), margins entire or with occasional tooth, (apex acute).
dense (elongated or not in fruit, 2.5–8.5(–10) cm).
congested, (elongated in fruit, greatly exceeding leaves).
sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate);
petals spatulate to narrowly oblanceolate, 7–10(–12) mm.
sepals lanceolate to oblong, 6–8 mm, (often keeled);
petals spatulate, 10–12 mm.
(divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels).
(spreading, straight or slightly curved), 8–12 mm.
didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave);
valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed;
replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute;
ovules 4–8 per ovary;
style 2–9 mm, (usually not exceeding sinus).
(erect), didymous, inflated, 10–20 × 10–20 mm, (papery or firm, basal sinus shallow to deep, sometimes barely notched, apical sinus deep, narrow, usually closed);
valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading (appearing fuzzy);
replum obovate to broadly oblong, not constricted, 3–4 mm, as wide as or wider than fruit, apex obtuse;
ovules (4–)8 per ovary;
style 7–9 mm.
slightly flattened, (ovate).
flattened.
Physaria newberryi
Physaria didymocarpa
Subspecies 2 (2 in the flora).
Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Subspecies 3 (3 in the flora).
The characters used to differentiate Physaria didymocarpa from P. saximontana (especially subsp. dentata) appear to be weak at best: whether there are 4 or 8 ovules per ovary and whether the silicle lacks a basal sinus or one is present. There appears to be intergradation in each of those characters. A traditional circumscription of these species is followed here. Further work is needed at both the species and subspecies level in these taxa.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Styles less than 4 mm, shorter than fruit sinuses. | subsp. newberryi |
1. Styles 5-9 mm, longer than fruit sinuses. | subsp. yesicola |
1. Trichomes spreading throughout; basal leaf blade margins dentate. | subsp. lanata |
1. Trichomes appressed (except fruits in subsp. didymocarpa); basal leaf blade margins repand to dentate, ± lyrate or, rarely, entire | → 2 |
2. Fruits moderately inflated, trichomes spreading. | subsp. didymocarpa |
2. Fruits highly inflated (papery), trichomes appressed. | subsp. lyrata |
CalFlora
WildflowerSearch
iNaturalist
USDA Plants Database
BC
WA
Flora NW
PNW Herbaria
