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Newberry twinpod, Newberry's twinpod

denseflower bladderpod, low bladderpod

Habit Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. Annuals or biennials; caudex simple or branched, (relatively small, cespitose); densely pubescent, trichomes (spreading, sessile or short-stalked), 5–7-rayed, rays distinct and simple, (tuberculate, finely tubercled with a U-shaped notch on one side).
Stems

several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm.

simple or few to several from base, erect or decumbent, (rarely branched, usually leafy), to 4 dm.

Basal leaves

(ascending to erect, petiole slender);

blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse).

blade lyrate-pinnatifid, 1–7 cm, margins entire or shallowly dentate.

Cauline leaves

blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire.

(sessile or shortly petiolate);

blade narrowly obovate to elliptic, 0.5–6 cm, margins entire, repand, or shallowly dentate.

Racemes

dense (elongated or not in fruit, 2.5–8.5(–10) cm).

dense, (elongated in fruit, often subtended by distal cauline leaves).

Flowers

sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate);

petals spatulate to narrowly oblanceolate, 7–10(–12) mm.

sepals elliptic, 3.7–7.2 mm, (lateral pair somewhat cucullate, median pair thickened apically);

petals (yellow to orange-yellow), obovate to obdeltate, (4.5–)7–10(–11) mm, (tapering to short claw, apex often emarginate).

Fruiting pedicels

(divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels).

(usually divaricate-spreading, straight or slightly curved, delicate, sometimes drooping, especially on herbarium specimens), 7–10 mm, (somewhat rigid).

Fruits

didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave);

valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed;

replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute;

ovules 4–8 per ovary;

style 2–9 mm, (usually not exceeding sinus).

(sessile or substipitate), globose or broadly obovate, not inflated, 4–6 mm, (smooth);

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules 8–16 per ovary;

style 2–5 mm.

Seeds

slightly flattened, (ovate).

flattened.

2n

= 14.

Physaria newberryi

Physaria densiflora

Phenology Flowering Mar–May.
Habitat Sandy, granitic, or calcareous soils, sandy ledges, limestone outcrops, rocky prairies, uplands
Elevation 30-400 m (100-1300 ft)
Distribution
from FNA
AZ; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Alyssum densiflorum (A. Gray) Kuntze (1891), not Desfontaines (1808) is an illegitimate name, sometimes found in synonymy with Physaria densiflora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles less than 4 mm, shorter than fruit sinuses.
subsp. newberryi
1. Styles 5-9 mm, longer than fruit sinuses.
subsp. yesicola
Source FNA vol. 7, p. 652. FNA vol. 7, p. 633.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. newberryi subsp. newberryi, P. newberryi subsp. yesicola
Synonyms P. didymocarpa var. newberryi Vesicaria densiflora, Lesquerella densiflora
Name authority A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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