Physaria newberryi |
Physaria cinerea |
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Newberry twinpod, Newberry's twinpod |
basin bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex branched, (woody); densely pubescent, trichomes (appressed, except spreading on pedicels and fruits, sessile or short-stalked), several-rayed, rays furcate or bifurcate, (strongly tuberculate throughout). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
few from base, erect, (stout), to 1.5 dm (sometimes greatly reduced). |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
blade suborbicular to rhombic or broadly elliptic, 1–4.5 cm, margins entire, (apex rounded to subacute). |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(proximal petiolate, distal subsessile); blade elliptic, 1–4 cm, (distal narrower), margins entire or remotely dentate. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
condensed, (subcorymbose to subumbellate, few-flowered). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals (persistent), broadly ovate or oblong to narrowly elliptic, 5.5–8(–9.5) mm, (lateral pair slightly cucullate, median pair thickened apically, cucullate, usually keeled); petals (orange to yellow), oblong to obovate, 8–11.5(–14.5) mm, (slightly narrowed to broad claw, margins sinuate, often retuse). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(horizontal or divaricate-ascending, straight or slightly curved), 5–15 mm, (stout and rigid). |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(sessile or substipitate), globose, ovoid, or suborbicular, compressed (with marginal and apical constriction), 4–7 mm; valves pubescent, trichomes contiguous or overlapping, often spreading; ovules 16–24 per ovary; style 2–4 mm. |
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Seeds | slightly flattened, (ovate). |
lenticular, ovate in outline. |
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2n | = 10. |
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Physaria newberryi |
Physaria cinerea |
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Phenology | Flowering Mar–May. | |||||
Habitat | Red soil, chiprock, gypsum or chalky knolls, limestone rubble | |||||
Elevation | 900-2200 m (3000-7200 ft) | |||||
Distribution |
AZ; NM; NV; UT
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AZ |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 631. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella cinerea | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||
Web links |