Physaria newberryi |
Physaria chambersii |
|||||
---|---|---|---|---|---|---|
Newberry twinpod, Newberry's twinpod |
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod |
|||||
Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex usually simple, sometimes branched, (thick, cespitose); (silvery) pubescent throughout, trichomes few-rayed, rays furcate, sometimes slightly fused at base, (umbonate, lightly tuberculate to nearly smooth). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm. |
||||
Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(petiole slender); blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate. |
||||
Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute). |
||||
Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
congested. |
||||
Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals narrowly lanceolate, 5–8(–9) mm; petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade). |
||||
Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(divaricate, slightly sigmoid), 8–15 mm. |
||||
Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded); valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent; replum oblong, as wide as or wider than fruits, apex obtuse; ovules 4–12 per ovary; style (4–)6–8 mm (exceeding sinus). |
||||
Seeds | slightly flattened, (ovate). |
flattened. |
||||
2n | = 8, 10, 16, 24. |
|||||
Physaria newberryi |
Physaria chambersii |
|||||
Phenology | Flowering Apr–Jul. | |||||
Habitat | Clay hillsides, limestone gravel, dolomite ridges, roadbanks, loose gravel, reddish clay, sagebrush and pinyon-juniper areas | |||||
Elevation | 1500-3200 m (4900-10500 ft) | |||||
Distribution |
AZ; NM; NV; UT
|
AZ; CA; ID; NV; OR; UT
|
||||
Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||
Key |
|
|||||
Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 631. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | P. chambersii var. canaani, P. chambersii var. sobolifera | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | Rollins: Rhodora 41: 403, plate 556, figs. 15–18. (1939) | ||||
Web links |