Physaria newberryi |
Physaria calcicola |
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Newberry twinpod, Newberry's twinpod |
Rocky Mountain bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; (compact); caudex branched; densely (silvery) pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays distinct, furcate or bifurcate, (umbonate, tuberculate and the center less so). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
several from base, erect or outer ones decumbent, (unbranched, stout, usually sparsely leaved), 1–3 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
blade linear, 2–7(–10) cm, margins entire, repand, or shallowly dentate. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(sessile); blade (erect), spatulate to linear, (1–)2–3(–4.5) cm, margins entire, sometimes involute, (apex acute or subacute). |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
dense, (exceeding basal leaves). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals ovate or oblong, (4.5–)5–6(–7) mm, (lateral pair subsaccate, cucullate, median pair thickened, cucullate apically); petals spatulate, 7–9(–11) mm (widened at base, slightly retuse). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(spreading, sharply sigmoid), 8–15 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(sessile or substipitate), ovate to oblong, not compressed at distal margins or apex, 5–9 mm; valves sparsely pubescent, trichomes appressed; ovules 4–8 per ovary; style 3–5 mm. |
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Seeds | slightly flattened, (ovate). |
flattened. |
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2n | = 16, ca. 20. |
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Physaria newberryi |
Physaria calcicola |
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Phenology | Flowering May–Jun. | |||||
Habitat | Shale bluffs, limestone hillsides, gypseous knolls and ravines, calcareous substrates, grasslands and pinyon-juniper communities | |||||
Elevation | 1400-2100 m (4600-6900 ft) | |||||
Distribution |
AZ; NM; NV; UT
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CO; NM
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 629. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella calcicola | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||
Web links |