Physaria newberryi |
Physaria aurea |
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Newberry twinpod, Newberry's twinpod |
golden bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Biennials or perennials; (short-lived); caudex branched; densely pubescent, trichomes (sessile or short-stalked, simple or not), 5–9-rayed, rays furcate, (fine, smooth or finely tuberculate). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
several from base, erect or outer ones decumbent or procumbent, (sometimes much-branched distally), to 6 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
blade obovate or rhombic, to ca. 2.5 cm, margins usually shallowly dentate, sometimes lyrate-pinnatifid. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(proximal shortly petiolate, distal sessile); blade obovate to rhombic or oblanceolate, 2–4(–6) cm, margins entire or shallowly and remotely dentate. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
usually dense, (several-flowered). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals ovate or oblong (tapering at base), 3.6–4.8(–5.3) mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals obovate to spatulate, 4.5–7.5 mm, (blade narrowed to broad claw, margins sinuate). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(strongly recurved), to 20 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(± pendent), ovoid, obcompressed, or globose, compressed, 4–6(–8) mm; valves (not retaining seeds after dehiscence), sparsely pubescent or glabrous, sparsely pubescent inside; replum as wide as or wider than fruit; ovules usually 4, rarely 6, per ovary; style 2.5–3.6 mm. |
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Seeds | slightly flattened, (ovate). |
flattened. |
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2n | = 14. |
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Physaria newberryi |
Physaria aurea |
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Phenology | Flowering Jun–Aug. | |||||
Habitat | Open sites and bare areas in rocky limestone soil in mountains, roadbanks, open woods | |||||
Elevation | 2000-2800 m (6600-9200 ft) | |||||
Distribution |
AZ; NM; NV; UT
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NM |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria aurea (known from the Jicarilla and Sacramento mountains) is similar to 35. P. gooddingii, which is found farther west in the mountains of Catron and Sierra counties, New Mexico, and Greenlee County, Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 628. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Lesquerella aurea | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Wooton) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) | ||||
Web links |