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Newberry twinpod, Newberry's twinpod

Arizona bladderpod

Habit Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. Perennials; caudex branched, (cespitose); densely (silvery gray) pubescent, trichomes (sessile or subsessile), (4-), 6-, or 8-rayed, rays fused at base, furcate or bifurcate, (slightly umbonate, tuberculate throughout).
Stems

several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm.

simple or few to several from base, erect, (unbranched, slender), 0.2–1(–1.5) dm.

Basal leaves

(ascending to erect, petiole slender);

blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse).

(densely tufted, not rosulate, reflexed in age);

blade obovate to oblanceolate, 0.7–2(–3) cm, margins usually entire, sometimes repand or shallowly dentate, (apex acute).

Cauline leaves

blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire.

similar to basal, becoming narrower distally, somewhat reflexed, (distal) blade linear or narrowly oblanceolate, 0.5–2.5(–5.5) cm.

Racemes

dense (elongated or not in fruit, 2.5–8.5(–10) cm).

dense, often subcorymbiform.

Flowers

sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate);

petals spatulate to narrowly oblanceolate, 7–10(–12) mm.

sepals (green or greenish yellow), ovate or broadly ovate, 3.5–6.5 mm, (lateral pair subsaccate, cucullate, median pair thickened, slightly cucullate apically);

petals (spreading), oblanceolate to obovate, 5.5–8(–10) mm, (claw erect).

Fruiting pedicels

(divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels).

(erect or divaricate-spreading, straight or slightly curved), (3–)5–10(–15) mm.

Fruits

didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave);

valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed;

replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute;

ovules 4–8 per ovary;

style 2–9 mm, (usually not exceeding sinus).

(sessile or substipitate), suborbicular to ovoid or ellipsoid, slightly inflated, 4–7 mm;

valves pubescent outside, trichomes substipitate, spreading, sometimes sparsely pubescent inside, trichomes sessile, smooth;

ovules 4–10(–16) per ovary;

style (0.5–)1–2(–4) mm (shorter than fruit).

Seeds

slightly flattened, (ovate).

flattened.

2n

= 10.

Physaria newberryi

Physaria arizonica

Phenology Flowering Apr–Jun.
Habitat Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine
Elevation 1000-2200 m (3300-7200 ft)
Distribution
from FNA
AZ; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The circumscription of Physaria arizonica here is quite broad and includes plants that have densely tufted basal leaves and relatively few or no cauline leaves; plants that are loosely tufted and have several cauline leaves; and plants that have a strongly branched caudex, leafy stems, and sterile shoots (var. andrusensis). Additional study is needed to understand the pattern of variation in this complex species; all of the characters given above vary considerably.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles less than 4 mm, shorter than fruit sinuses.
subsp. newberryi
1. Styles 5-9 mm, longer than fruit sinuses.
subsp. yesicola
Source FNA vol. 7, p. 652. FNA vol. 7, p. 627.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. newberryi subsp. newberryi, P. newberryi subsp. yesicola
Synonyms P. didymocarpa var. newberryi Lesquerella arizonica, Lesquerella arizonica var. nudicaulis, P. arizonica var. andrusensis
Name authority A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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