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Newberry twinpod, Newberry's twinpod

Great Plains bladderpod

Habit Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. Perennials or, rarely, annuals; caudex simple or branched; ± densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (usually spreading), distinct or slightly fused at base, furcate or bifurcate, (tuberculate).
Stems

several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm.

simple or few from base, prostrate or straggling to erect, (sometimes purplish, usually unbranched), (0.5–)1–2(–3) dm.

Basal leaves

(ascending to erect, petiole slender);

blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse).

blade oblanceolate, 1.5–5(–7) cm, margins entire or shallowly dentate, (flat).

Cauline leaves

blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire.

blade elliptic to linear, (0.5–)1–2.5(–3) cm, margins usually entire.

Racemes

dense (elongated or not in fruit, 2.5–8.5(–10) cm).

(secund), dense, (elongated in fruit).

Flowers

sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate);

petals spatulate to narrowly oblanceolate, 7–10(–12) mm.

sepals elliptic or oblong, 4–6(–7) mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals (often red or lavender when dried), obovate, 6–8.5(–9.5) mm, (narrowing to broad claw).

Fruiting pedicels

(divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels).

(usually sharply recurved, sometimes divaricate-spreading or nearly horizontal), 5–15(–20) mm, (stout).

Fruits

didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave);

valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed;

replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute;

ovules 4–8 per ovary;

style 2–9 mm, (usually not exceeding sinus).

subglobose, obovoid, or broadly ellipsoid, slightly inflated, (3.5–)4–5.5(–6.5) mm;

valves densely pubescent outside, trichomes spreading or closely appressed, rarely sparsely pubescent inside;

ovules (4–)8(–10) per ovary;

style (slender), 3–5.5(–6.5) mm.

Seeds

slightly flattened, (ovate).

slightly flattened.

Physaria newberryi

Physaria arenosa

Distribution
from FNA
AZ; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; MT; ND; NE; SD; WY; AB; MB; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles less than 4 mm, shorter than fruit sinuses.
subsp. newberryi
1. Styles 5-9 mm, longer than fruit sinuses.
subsp. yesicola
1. Perennials or, rarely, annuals, short-lived; fruit valves: trichomes spreading.
subsp. arenosa
1. Perennials, long-lived; fruit valves: trichomes closely appressed.
subsp. argillosa
Source FNA vol. 7, p. 652. FNA vol. 7, p. 626.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. newberryi subsp. newberryi, P. newberryi subsp. yesicola
P. arenosa subsp. arenosa, P. arenosa subsp. argillosa
Synonyms P. didymocarpa var. newberryi Vesicaria arenosa, Lesquerella arenosa, Lesquerella argentea var. arenosa, Lesquerella ludoviciana var. arenosa
Name authority A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) (Richardson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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