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Newberry twinpod, Newberry's twinpod

arctic bladderpod

Habit Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. Perennials; caudex simple or branched, (woody, cespitose); ± densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays distinct, furcate or bifurcate, (somewhat umbonate, finely tuberculate to ± smooth).
Stems

several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm.

simple or few to several from base, erect to spreading or prostrate, 0.5–2(–3) dm.

Basal leaves

(ascending to erect, petiole slender);

blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse).

(usually ± rosulate);

blade obovate to oblanceolate, (1–)2–6(–15) cm, margins entire.

Cauline leaves

blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire.

(sessile or shortly petiolate);

blade oblanceolate or lingulate, 0.5–1.5(–3) cm, margins entire.

Racemes

dense (elongated or not in fruit, 2.5–8.5(–10) cm).

loose.

Flowers

sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate);

petals spatulate to narrowly oblanceolate, 7–10(–12) mm.

sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate);

petals spatulate, 5–6(–7) mm, (blade gradually narrowed to claw).

Fruiting pedicels

(divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels).

(erect to divaricate or ascending), (5–)10–20(–40) mm, (stout).

Fruits

didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave);

valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed;

replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute;

ovules 4–8 per ovary;

style 2–9 mm, (usually not exceeding sinus).

subglobose to ellipsoid, uncompressed, 4–6(–9) mm;

valves (not retaining seeds after dehiscence), glabrous or sparsely pubescent outside, trichomes sessile;

replum as wide as or wider than fruit;

ovules (8–)10–14(–16) per ovary;

style 1–2.5(–4) mm.

Seeds

slightly flattened, (ovate).

plump.

2n

= 60.

Physaria newberryi

Physaria arctica

Phenology Flowering May–Aug.
Habitat Typically on sand and gravel from calcareous bedrock, river bars and terraces, cliff ledges, scree and talus slopes, often growing after disturbance
Elevation 0-1800 m (0-5900 ft)
Distribution
from FNA
AZ; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; circumarctic (except n Europe, ne Russia)
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles less than 4 mm, shorter than fruit sinuses.
subsp. newberryi
1. Styles 5-9 mm, longer than fruit sinuses.
subsp. yesicola
Source FNA vol. 7, p. 652. FNA vol. 7, p. 626.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. newberryi subsp. newberryi, P. newberryi subsp. yesicola
Synonyms P. didymocarpa var. newberryi Alyssum arcticum, Lesquerella arctica, Lesquerella arctica subsp. purshii, Vesicaria arctica, Vesicaria leiocarpa
Name authority A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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