Physaria newberryi |
Physaria arctica |
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Newberry twinpod, Newberry's twinpod |
arctic bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex simple or branched, (woody, cespitose); ± densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays distinct, furcate or bifurcate, (somewhat umbonate, finely tuberculate to ± smooth). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
simple or few to several from base, erect to spreading or prostrate, 0.5–2(–3) dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(usually ± rosulate); blade obovate to oblanceolate, (1–)2–6(–15) cm, margins entire. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(sessile or shortly petiolate); blade oblanceolate or lingulate, 0.5–1.5(–3) cm, margins entire. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
loose. |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals spatulate, 5–6(–7) mm, (blade gradually narrowed to claw). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(erect to divaricate or ascending), (5–)10–20(–40) mm, (stout). |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
subglobose to ellipsoid, uncompressed, 4–6(–9) mm; valves (not retaining seeds after dehiscence), glabrous or sparsely pubescent outside, trichomes sessile; replum as wide as or wider than fruit; ovules (8–)10–14(–16) per ovary; style 1–2.5(–4) mm. |
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Seeds | slightly flattened, (ovate). |
plump. |
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2n | = 60. |
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Physaria newberryi |
Physaria arctica |
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Phenology | Flowering May–Aug. | |||||
Habitat | Typically on sand and gravel from calcareous bedrock, river bars and terraces, cliff ledges, scree and talus slopes, often growing after disturbance | |||||
Elevation | 0-1800 m (0-5900 ft) | |||||
Distribution |
AZ; NM; NV; UT
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AK; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; circumarctic (except n Europe, ne Russia) |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 626. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Alyssum arcticum, Lesquerella arctica, Lesquerella arctica subsp. purshii, Vesicaria arctica, Vesicaria leiocarpa | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | ||||
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