Physaria newberryi |
Physaria angustifolia |
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Newberry twinpod, Newberry's twinpod |
threadleaf bladderpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Annuals; with a fine taproot; ± densely pubescent, trichomes several-rayed, rays distinct or fused at base, bifurcate, (prominently tuberculate throughout). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
simple or few to several from base, erect, (sometimes branched), to 4 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
blade elliptic to rhombic, 3–8 cm, (base narrowing gradually to petiole), margins entire, repand, coarsely toothed, or pinnatifid. |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
(proximal often shortly petiolate, distal sessile); blade linear or narrowly obovate, 1.5–6(–10) cm, margins entire, repand, or shallowly toothed. |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
usually loose. |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals elliptic or ovate, 4–6 mm, (lateral pair usually subsaccate); petals obovate to obdeltate, 6–10 mm, (apex often emarginate). |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(usually divaricate, sometimes horizontal, straight or slightly curved), 8–20 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
not didymous, ± globose, slightly inflated, 4–6 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4 per ovary; style 2–3.5 mm; (stigma expanded). |
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Seeds | slightly flattened, (ovate). |
flattened, (margined). |
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2n | = 10. |
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Physaria newberryi |
Physaria angustifolia |
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Phenology | Flowering Apr(-May). | |||||
Habitat | Shallow limestone-derived soils, sometimes spreading to disturbed sites | |||||
Elevation | 90-300 m (300-1000 ft) | |||||
Distribution |
AZ; NM; NV; UT
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OK; TX |
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 625. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | Vesicaria angustifolia, Lesquerella angustifolia, Lesquerella longifolia | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | (Nuttall ex Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | ||||
Web links |