Physaria newberryi |
Physaria acutifolia |
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Newberry twinpod, Newberry's twinpod |
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
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Habit | Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | ||||
Stems | several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm. |
several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
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Basal leaves | (ascending to erect, petiole slender); blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse). |
(petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
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Cauline leaves | blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire. |
blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
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Racemes | dense (elongated or not in fruit, 2.5–8.5(–10) cm). |
loose, (elongated in fruit). |
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Flowers | sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate); petals spatulate to narrowly oblanceolate, 7–10(–12) mm. |
sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
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Fruiting pedicels | (divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels). |
(divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
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Fruits | didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave); valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed; replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute; ovules 4–8 per ovary; style 2–9 mm, (usually not exceeding sinus). |
(erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
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Seeds | slightly flattened, (ovate). |
(dark brown), flattened, (2–3 mm). |
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2n | = 10, 16, 24. |
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Physaria newberryi |
Physaria acutifolia |
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Phenology | Flowering May–Jun(-Jul). | |||||
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | |||||
Elevation | 1500-3500 m (4900-11500 ft) | |||||
Distribution |
AZ; NM; NV; UT
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AZ; CO; ID; MT; NM; NV; SD; UT; WY
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Discussion | Subspecies 2 (2 in the flora). Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 652. | FNA vol. 7, p. 624. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. didymocarpa var. newberryi | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | ||||
Name authority | A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | ||||
Web links |