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Newberry twinpod, Newberry's twinpod

Habit Perennials; caudex simple or branched, (branches often covered with persistent leaf bases, cespitose); densely (silvery) pubescent, trichomes rays fused at least 1/2 their length. Annuals, biennials, perennials, or subshrubs; eglandular.
Stems

several from base, ascending to erect (arising laterally, unbranched), 0.5–1(–2.5) dm.

Basal leaves

(ascending to erect, petiole slender);

blade oblanceolate to obovate, 3–8 cm, (base tapering to petiole), margins incised or dentate with broad teeth, (apex acute to obtuse).

Cauline leaves

blade linear-oblanceolate to oblanceolate, 1–2 cm, margins entire.

petiolate, sessile, or subsessile;

blade base usually not auriculate (except Paysonia), margins entire, dentate, or sinuate.

Racemes

dense (elongated or not in fruit, 2.5–8.5(–10) cm).

ebracteate, often elongated in fruit.

Flowers

sepals (greenish yellow), lanceolate, 6–8.5 mm, (saccate and cucullate);

petals spatulate to narrowly oblanceolate, 7–10(–12) mm.

actinomorphic;

sepals erect, spreading, ascending, or reflexed, lateral pair seldom saccate basally;

petals white, yellow, lavender, purple, violet, orange, or brown [pink], claw present, often distinct;

filaments unappendaged, not winged;

pollen (3 or) 4–11-colpate.

Fruiting pedicels

(divaricate, straight), 5–11(–15) mm, (rigid, fruits not pendent on arching pedicels).

Fruits

didymous, sides curved and angular, highly inflated, 6–16 × 8–12 mm, (papery, apical sinus broad and concave);

valves (retaining seeds after dehiscence, distinctly 2-keeled on side away from replum), pubescent, trichomes appressed;

replum linear to linear-lanceolate, as wide as or wider than fruit, apex acute;

ovules 4–8 per ovary;

style 2–9 mm, (usually not exceeding sinus).

silicles or siliques, dehiscent, unsegmented, terete, latiseptate, or angustiseptate;

ovules 2–100 per ovary;

style usually distinct;

stigma entire or strongly 2-lobed.

Seeds

slightly flattened, (ovate).

biseriate, uniseriate, or aseriate;

cotyledons accumbent or incumbent.

Trichomes

usually short-stalked, subsessile, or sessile, sometimes long-stalked, stellate, scalelike, subdendritic, or forked, sometimes mixed with simple ones.

Physaria newberryi

Brassicaceae tribe Physarieae

Distribution
from FNA
AZ; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
North America; Mexico; South America; Asia (ne Russia)
Discussion

Subspecies 2 (2 in the flora).

Physaria newberryi, with its unusual fruits, can be confused with 15. P. chambersii. In P. chambersii, the sides of the fruit are flat, the style always exceeds the top, or shoulders, of the fruit, and shoulders form an angle that does not curve in toward the style. In P. newberryi, the sides of the fruit are concave, the styles are shorter than shoulders of the silicle (except in subsp. yesicola), and shoulders of the silicle form a curved, inward arching crown on the fruit.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera 7, species ca. 130 (7 genera, 105 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Styles less than 4 mm, shorter than fruit sinuses.
subsp. newberryi
1. Styles 5-9 mm, longer than fruit sinuses.
subsp. yesicola
Source FNA vol. 7, p. 652. FNA vol. 7, p. 604.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. newberryi subsp. newberryi, P. newberryi subsp. yesicola
Synonyms P. didymocarpa var. newberryi
Name authority A. Gray: in J. C. Ives, Rep. Colorado R. 4: 6. (1861) B. L. Robinson: in A. Gray et al., Syn. Fl. N. Amer. 1(1,1): 100. (1895)
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