Physaria multiceps |
Physaria hitchcockii |
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manyhead bladderpod |
Hitchcock's bladderpod |
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Habit | Perennials; caudex (buried), branched, (not thickened); densely pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays furcate or bifurcate, (umbonate, rough to finely tuberculate throughout). | Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate). | ||||||||
Stems | several from base, prostrate, (slender, sparsely pubescent), 0.5–2 dm. |
few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm. |
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Basal leaves | blade obovate to narrowly elliptic, 1.5–6 cm, margins usually entire, rarely shallowly dentate, (surfaces densely pubescent, often silvery). |
(petiole and blade differentiated or not); blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire. |
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Cauline leaves | blade oblanceolate to spatulate, 0.5–1 cm, margins entire, (surfaces often sparsely pubescent). |
similar to basal, smaller. |
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Racemes | (narrow), loose, (elongated in fruit). |
dense. |
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Flowers | sepals (greenish brown, sometimes magenta), linear or elliptic, 4.3–6(–7.5) mm, (median pair thickened apically, cucullate); petals (frequently pink or magenta in distal 1/3–1/2), spatulate to oblanceolate, 6–10(–12) mm, (claw undifferentiated from blade). |
sepals narrowly lanceolate to lanceolate, 2.8–6 mm; petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (ascending to somewhat spreading, straight to slightly curved), 4–8(–12) mm. |
(ascending, straight or slightly curved), 2–6 mm. |
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Fruits | broadly ovoid to suborbicular, inflated, (terete or, often, slightly angustiseptate), 3–6mm; valves sparsely pubescent; ovules usually 4, rarely 6–8 per ovary; style 3–6.5 mm. |
(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute); valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8 per ovary; style 1.7–6 mm. |
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Seeds | plump. |
flattened. |
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Physaria multiceps |
Physaria hitchcockii |
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Phenology | Flowering May–Jul. | |||||||||
Habitat | Douglas-fir or spruce woodlands, limestone ridges, damp open slopes, soil pockets among rocks, crevices of rocks, decomposed calcareous rocks | |||||||||
Elevation | 2400-2900 m (7900-9500 ft) | |||||||||
Distribution |
ID; UT; WY |
NV; UT |
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Discussion | Subspecies 3 (3 in the flora). The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 651. | FNA vol. 7, p. 643. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Lesquerella multiceps | Lesquerella hitchcockii | ||||||||
Name authority | (Maguire) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | ||||||||
Web links |