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manyhead bladderpod

limestone glade or Missouri bladderpod

Habit Perennials; caudex (buried), branched, (not thickened); densely pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays furcate or bifurcate, (umbonate, rough to finely tuberculate throughout). Annuals; with a fine taproot; densely pubescent, trichomes (sessile), usually 4-rayed, rays forked, rarely simple or tripartite, (finely tuberculate).
Stems

several from base, prostrate, (slender, sparsely pubescent), 0.5–2 dm.

several from base, (slender), erect or outer decumbent, (usually branched, branches filiform, bud clusters of growing plants drooping), to 2.5 dm.

Basal leaves

blade obovate to narrowly elliptic, 1.5–6 cm, margins usually entire, rarely shallowly dentate, (surfaces densely pubescent, often silvery).

blade orbicular to broadly spatulate, 1–2.4 cm, margins entire or sinuate.

Cauline leaves

blade oblanceolate to spatulate, 0.5–1 cm, margins entire, (surfaces often sparsely pubescent).

(proximal often petiolate, distal sessile);

similar to basal, blade spatulate to oblanceolate or (distal) linear, (base cuneate), margins entire or sinuate.

Racemes

(narrow), loose, (elongated in fruit).

loose.

Flowers

sepals (greenish brown, sometimes magenta), linear or elliptic, 4.3–6(–7.5) mm, (median pair thickened apically, cucullate);

petals (frequently pink or magenta in distal 1/3–1/2), spatulate to oblanceolate, 6–10(–12) mm, (claw undifferentiated from blade).

sepals oblong or elliptic, 2.5–4.6 mm, (median pair slightly thickened apically);

petals (pale yellow), spatulate to obovate, 5–9 mm, (apex ± emarginate).

Fruiting pedicels

(ascending to somewhat spreading, straight to slightly curved), 4–8(–12) mm.

(usually divaricate-ascending, straight or slightly curved), 7–11 mm.

Fruits

broadly ovoid to suborbicular, inflated, (terete or, often, slightly angustiseptate), 3–6mm;

valves sparsely pubescent;

ovules usually 4, rarely 6–8 per ovary;

style 3–6.5 mm.

(sessile or shortly stipitate), globose, not inflated, 3–4 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4 per ovary;

style 3–5 mm.

Seeds

plump.

flattened.

2n

= 14.

Physaria multiceps

Physaria filiformis

Phenology Flowering May–Jul. Flowering Apr–May.
Habitat Douglas-fir or spruce woodlands, limestone ridges, damp open slopes, soil pockets among rocks, crevices of rocks, decomposed calcareous rocks Limestone, dolomite, and shale, sparsely vegetated or barren areas, cedar glades, old pastures, along roadsides
Elevation 2400-2900 m (7900-9500 ft) 200-300 m (700-1000 ft)
Distribution
from FNA
ID; UT; WY
[BONAP county map]
from FNA
AL; AR; MO
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 651. FNA vol. 7, p. 637.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella multiceps Lesquerella filiformis
Name authority (Maguire) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
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