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manyhead bladderpod

tuft twinpod

Habit Perennials; caudex (buried), branched, (not thickened); densely pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays furcate or bifurcate, (umbonate, rough to finely tuberculate throughout). Perennials; caudex usually simple, rarely branched, (enlarged with persistent leaf bases, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays typically furcate, (fused at base, arms slender, tuberculate throughout).
Stems

several from base, prostrate, (slender, sparsely pubescent), 0.5–2 dm.

several from base, decumbent to ascending, (arising laterally beneath a dense rosette), less than 0.1 dm.

Basal leaves

blade obovate to narrowly elliptic, 1.5–6 cm, margins usually entire, rarely shallowly dentate, (surfaces densely pubescent, often silvery).

(petiole slender);

blade (horizontal on the ground), obovate, 0.5–1.5 cm (width 4–8 mm, base tapering abruptly to petiole), margins entire, (apex usually acute, surfaces silvery from a dense incrustation of appressed, stellate trichomes).

Cauline leaves

blade oblanceolate to spatulate, 0.5–1 cm, margins entire, (surfaces often sparsely pubescent).

blade oblanceolate, 0.5–1 cm (width 2–3 mm), margins entire, (surfaces densely stellate pubescent).

Racemes

(narrow), loose, (elongated in fruit).

congested, (subumbellate, often almost sessile, barely exceeding basal leaves).

Flowers

sepals (greenish brown, sometimes magenta), linear or elliptic, 4.3–6(–7.5) mm, (median pair thickened apically, cucullate);

petals (frequently pink or magenta in distal 1/3–1/2), spatulate to oblanceolate, 6–10(–12) mm, (claw undifferentiated from blade).

sepals (yellowish green), narrowly lanceolate, 4–5 mm;

petals (erect), oblanceolate, 6–7 mm, (claw weakly differentiated from blade).

Fruiting pedicels

(ascending to somewhat spreading, straight to slightly curved), 4–8(–12) mm.

(divaricate, straight), 5–10 mm.

Fruits

broadly ovoid to suborbicular, inflated, (terete or, often, slightly angustiseptate), 3–6mm;

valves sparsely pubescent;

ovules usually 4, rarely 6–8 per ovary;

style 3–6.5 mm.

didymous, ovate, inflated, 4.8–6 × 6–10 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), pubescent, trichomes loosely spreading;

replum obovate, 3–4 mm, as wide as or wider than fruit;

ovules 8 per ovary (2–4 abortive);

style 4–6 mm.

Seeds

plump.

flattened.

Physaria multiceps

Physaria condensata

Phenology Flowering May–Jul. Flowering May–Jun.
Habitat Douglas-fir or spruce woodlands, limestone ridges, damp open slopes, soil pockets among rocks, crevices of rocks, decomposed calcareous rocks Calcareous knolls and ridges, clay banks, limey slopes, shaley hills, clay patches
Elevation 2400-2900 m (7900-9500 ft) 1800-2400 m (5900-7900 ft)
Distribution
from FNA
ID; UT; WY
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 651. FNA vol. 7, p. 631.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella multiceps
Name authority (Maguire) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) Rollins: Rhodora 41: 407, plate 556, figs. 1, 9, 10. (1939)
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