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manyhead bladderpod

Physaria alpestris

alpine twin-pod, Washington bladder-pod, Washington twin-pod

Habit Perennials; caudex (buried), branched, (not thickened); densely pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays furcate or bifurcate, (umbonate, rough to finely tuberculate throughout). Perennials; caudex usually simple, rarely branched, (cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays (1- or) 2-bifurcate, (low-umbonate, tubercles relatively few, small).
Stems

several from base, prostrate, (slender, sparsely pubescent), 0.5–2 dm.

several from base, decumbent to ascending, (unbranched), 0.5–1.5 dm.

Basal leaves

blade obovate to narrowly elliptic, 1.5–6 cm, margins usually entire, rarely shallowly dentate, (surfaces densely pubescent, often silvery).

(petiole slender);

blade obovate, 3–5 cm (width 10–20 mm, base tapering abruptly to petiole), margins entire, (apex rarely slightly acute).

Cauline leaves

blade oblanceolate to spatulate, 0.5–1 cm, margins entire, (surfaces often sparsely pubescent).

blade oblanceolate, 0.5–1.5 cm (width 3–5 mm), margins entire.

Racemes

(narrow), loose, (elongated in fruit).

subcorymbose.

Flowers

sepals (greenish brown, sometimes magenta), linear or elliptic, 4.3–6(–7.5) mm, (median pair thickened apically, cucullate);

petals (frequently pink or magenta in distal 1/3–1/2), spatulate to oblanceolate, 6–10(–12) mm, (claw undifferentiated from blade).

sepals oblong, 8–10 mm;

petals spatulate, 12–14 mm.

Fruiting pedicels

(ascending to somewhat spreading, straight to slightly curved), 4–8(–12) mm.

(divaricate, straight), 5–10 mm.

Fruits

broadly ovoid to suborbicular, inflated, (terete or, often, slightly angustiseptate), 3–6mm;

valves sparsely pubescent;

ovules usually 4, rarely 6–8 per ovary;

style 3–6.5 mm.

didymous, mostly highly inflated (strongly flattened at least in 1/2 toward replum), 14–18 × 14–18 mm, (papery, basal sinus slightly notched, apical open, shallow);

valves (retaining seeds after dehiscence), evenly pubescent;

replum lanceolate, 7–10 mm, width 1.5–2.5 mm, as wide as or wider than fruit, apex acute to acuminate;

ovules 8–10 per ovary;

style 5–7 mm.

Seeds

plump.

flattened, (2–3 mm).

2n

= 48–52, 52, 64, 67–70.

Physaria multiceps

Physaria alpestris

Phenology Flowering May–Jul. Flowering May–Jun.
Habitat Douglas-fir or spruce woodlands, limestone ridges, damp open slopes, soil pockets among rocks, crevices of rocks, decomposed calcareous rocks Alpine scree, rocky ridges, talus slopes, volcanic sands and gravel, serpentine gravel, granitic slopes, mountain shrub, subalpine fir, and whitebark pine communities
Elevation 2400-2900 m (7900-9500 ft) (700-)1300-2400 m ((2300-)4300-7900 ft)
Distribution
from FNA
ID; UT; WY
[BONAP county map]
from FNA
WA
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 7, p. 651. FNA vol. 7, p. 624.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella multiceps Lesquerella alpestris
Name authority (Maguire) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) Suksdorf: W. Amer. Sci. 15: 58. (1906)
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