Physaria montana |
Physaria kingii |
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mountain bladderpod |
King bladderpod, King's bladder-pod |
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Habit | Perennials; caudex simple or branched, (often enlarged); densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays furcate or bifurcate, (tuberculate throughout). | Perennials; caudex usually simple, sometimes branched, (not thickened); usually densely pubescent, trichomes (sessile or short-stalked), 3–7-rayed, rays distinct or slightly fused at base, typically furcate near base, bifurcate or 3-partite, (not to slightly umbonate, smooth or moderately to strongly tuberculate). | ||||||||||||||||||||||||
Stems | simple or several from base, prostrate to erect, 0.5–2(–3.5) dm. |
few to several from base, prostrate to decumbent or erect, 0.5–2(–4) dm. |
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Basal leaves | blade suborbicular or obovate to elliptic, (1–)2–5(–7) cm, margins entire, sinuate, or shallowly dentate. |
blade suborbicular to narrowly or broadly oblanceolate to broadly elliptic or rhombic, (1.2–)2–6(–8) cm, (base usually abruptly narrowed to petiole), margins entire, sinuate, or lobed. |
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Cauline leaves | (often secund, proximal shortly petiolate, distal sessile); blade linear to obovate or rhombic, 1–2.5(–4) cm, margins entire or shallowly dentate. |
(proximal petiolate, distal sessile); blade obovate or elliptic to spatulate, 0.5–2 cm, margins entire. |
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Racemes | dense, compact, (usually elongated in fruit). |
(usually not secund), dense, (sometimes elongated in fruit). |
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Flowers | sepals elliptic, 5–8.5 mm, (lateral pair boat-shaped, saccate, median pair thickened apically, cucullate); petals (yellow to orange, sometimes fading purplish), narrowly spatulate or obovate, (6–)7.5–12 mm, (claw undifferentiated from blade, or gradually narrowed to claw, slightly expanded basally). |
sepals lanceolate, 4–6(–7) mm; petals (yellow, cream-yellow, cream-white, or white), obovate to oblanceolate, 6–13 mm, (claw weakly differentiated from blade). |
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Fruiting pedicels | (usually sharply sigmoid, rarely nearly divaricate-spreading and straight), 5–15(–20) mm, (stout). |
(erect to divaricate-ascending or recurved, erect in distal 1/3, usually sigmoid, sometimes straight or slightly curved), 4.5–10(–15) mm. |
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Fruits | (erect), ellipsoid or ovoid, not or slightly obcompressed, (apex not compressed), (6–)7–12 mm; valves densely pubescent, sometimes sparsely pubescent inside; ovules (8–)12–20(–24) per ovary; style 3–7 mm, (sometimes pubescent). |
(sessile or substipitate), subglobose, obovoid, or ellipsoid, compressed (sometimes slightly angustiseptate), 3–9 mm, (rigid, apex truncate, retuse, or rounded-acute); valves sparsely or densely pubescent, sometimes sparsely pubescent inside; (septum sometimes fenestrate, perforate, or obsolete); ovules 4–16 per ovary; style 1–9 mm. |
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Seeds | flattened. |
flattened (sometimes slightly). |
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2n | = 10. |
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Physaria montana |
Physaria kingii |
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Phenology | Flowering Apr–Jun(-Aug). | |||||||||||||||||||||||||
Habitat | Banks, rock outcrops, stony slopes and benchlands, from plains into mountains, in sagebrush, open scrub oak, pinyon-juniper woodland, ponderosa pine, Douglas fir on granitic, often gravelly, non-calcareous soils, rarely on calcareous soils | |||||||||||||||||||||||||
Elevation | 1000-3300 m (3300-10800 ft) | |||||||||||||||||||||||||
Distribution |
AZ; CO; NE; NM; SD; WY
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AZ; CA; ID; NV; OR; UT; nw Mexico
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Discussion | Physaria montana is a rather variable species that in southwestern Colorado morphologically approaches P. rectipes and in eastern Wyoming approaches P. curvipes; it is unusual in the genus for its frequent presence on igneous, non-calcareous soils. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 7 (7 in the flora). The Physaria kingii complex is in need of further study. It is widespread in the western United States, mostly in montane environments. This treatment recognizes a highly variable species with generally well-marked, geographically coherent subspecies. Hybridization may be involved in some of the subspecies, especially in subsp. kaibabensis, where molecular data indicate intra-individual genetic variation (pers. obs.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 650. | FNA vol. 7, p. 645. | ||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||||||||||||||
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Synonyms | Vesicaria montana, Alyssum grayanum, Lesquerella montana, Lesquerella montana var. suffruticosa, Lesquerella rosulata | Vesicaria kingii, Lesquerella kingii | ||||||||||||||||||||||||
Name authority | (A. Gray) Greene: Fl. Francisc., 249. (1891) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | ||||||||||||||||||||||||
Web links |