Physaria montana |
Physaria arizonica |
|
---|---|---|
mountain bladderpod |
Arizona bladderpod |
|
Habit | Perennials; caudex simple or branched, (often enlarged); densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays furcate or bifurcate, (tuberculate throughout). | Perennials; caudex branched, (cespitose); densely (silvery gray) pubescent, trichomes (sessile or subsessile), (4-), 6-, or 8-rayed, rays fused at base, furcate or bifurcate, (slightly umbonate, tuberculate throughout). |
Stems | simple or several from base, prostrate to erect, 0.5–2(–3.5) dm. |
simple or few to several from base, erect, (unbranched, slender), 0.2–1(–1.5) dm. |
Basal leaves | blade suborbicular or obovate to elliptic, (1–)2–5(–7) cm, margins entire, sinuate, or shallowly dentate. |
(densely tufted, not rosulate, reflexed in age); blade obovate to oblanceolate, 0.7–2(–3) cm, margins usually entire, sometimes repand or shallowly dentate, (apex acute). |
Cauline leaves | (often secund, proximal shortly petiolate, distal sessile); blade linear to obovate or rhombic, 1–2.5(–4) cm, margins entire or shallowly dentate. |
similar to basal, becoming narrower distally, somewhat reflexed, (distal) blade linear or narrowly oblanceolate, 0.5–2.5(–5.5) cm. |
Racemes | dense, compact, (usually elongated in fruit). |
dense, often subcorymbiform. |
Flowers | sepals elliptic, 5–8.5 mm, (lateral pair boat-shaped, saccate, median pair thickened apically, cucullate); petals (yellow to orange, sometimes fading purplish), narrowly spatulate or obovate, (6–)7.5–12 mm, (claw undifferentiated from blade, or gradually narrowed to claw, slightly expanded basally). |
sepals (green or greenish yellow), ovate or broadly ovate, 3.5–6.5 mm, (lateral pair subsaccate, cucullate, median pair thickened, slightly cucullate apically); petals (spreading), oblanceolate to obovate, 5.5–8(–10) mm, (claw erect). |
Fruiting pedicels | (usually sharply sigmoid, rarely nearly divaricate-spreading and straight), 5–15(–20) mm, (stout). |
(erect or divaricate-spreading, straight or slightly curved), (3–)5–10(–15) mm. |
Fruits | (erect), ellipsoid or ovoid, not or slightly obcompressed, (apex not compressed), (6–)7–12 mm; valves densely pubescent, sometimes sparsely pubescent inside; ovules (8–)12–20(–24) per ovary; style 3–7 mm, (sometimes pubescent). |
(sessile or substipitate), suborbicular to ovoid or ellipsoid, slightly inflated, 4–7 mm; valves pubescent outside, trichomes substipitate, spreading, sometimes sparsely pubescent inside, trichomes sessile, smooth; ovules 4–10(–16) per ovary; style (0.5–)1–2(–4) mm (shorter than fruit). |
Seeds | flattened. |
flattened. |
2n | = 10. |
= 10. |
Physaria montana |
Physaria arizonica |
|
Phenology | Flowering Apr–Jun(-Aug). | Flowering Apr–Jun. |
Habitat | Banks, rock outcrops, stony slopes and benchlands, from plains into mountains, in sagebrush, open scrub oak, pinyon-juniper woodland, ponderosa pine, Douglas fir on granitic, often gravelly, non-calcareous soils, rarely on calcareous soils | Sandy and gravelly soils, limey knolls or limestone chip, often in open stands of sagebrush-pinyon, pinyon-juniper, Gambel oak and sometimes ponderosa pine |
Elevation | 1000-3300 m (3300-10800 ft) | 1000-2200 m (3300-7200 ft) |
Distribution |
AZ; CO; NE; NM; SD; WY
|
AZ; UT
|
Discussion | Physaria montana is a rather variable species that in southwestern Colorado morphologically approaches P. rectipes and in eastern Wyoming approaches P. curvipes; it is unusual in the genus for its frequent presence on igneous, non-calcareous soils. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The circumscription of Physaria arizonica here is quite broad and includes plants that have densely tufted basal leaves and relatively few or no cauline leaves; plants that are loosely tufted and have several cauline leaves; and plants that have a strongly branched caudex, leafy stems, and sterile shoots (var. andrusensis). Additional study is needed to understand the pattern of variation in this complex species; all of the characters given above vary considerably. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 650. | FNA vol. 7, p. 627. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria montana, Alyssum grayanum, Lesquerella montana, Lesquerella montana var. suffruticosa, Lesquerella rosulata | Lesquerella arizonica, Lesquerella arizonica var. nudicaulis, P. arizonica var. andrusensis |
Name authority | (A. Gray) Greene: Fl. Francisc., 249. (1891) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
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