Physaria montana |
Physaria acutifolia |
|
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mountain bladderpod |
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
|
Habit | Perennials; caudex simple or branched, (often enlarged); densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays furcate or bifurcate, (tuberculate throughout). | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). |
Stems | simple or several from base, prostrate to erect, 0.5–2(–3.5) dm. |
several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
Basal leaves | blade suborbicular or obovate to elliptic, (1–)2–5(–7) cm, margins entire, sinuate, or shallowly dentate. |
(petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
Cauline leaves | (often secund, proximal shortly petiolate, distal sessile); blade linear to obovate or rhombic, 1–2.5(–4) cm, margins entire or shallowly dentate. |
blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
Racemes | dense, compact, (usually elongated in fruit). |
loose, (elongated in fruit). |
Flowers | sepals elliptic, 5–8.5 mm, (lateral pair boat-shaped, saccate, median pair thickened apically, cucullate); petals (yellow to orange, sometimes fading purplish), narrowly spatulate or obovate, (6–)7.5–12 mm, (claw undifferentiated from blade, or gradually narrowed to claw, slightly expanded basally). |
sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
Fruiting pedicels | (usually sharply sigmoid, rarely nearly divaricate-spreading and straight), 5–15(–20) mm, (stout). |
(divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
Fruits | (erect), ellipsoid or ovoid, not or slightly obcompressed, (apex not compressed), (6–)7–12 mm; valves densely pubescent, sometimes sparsely pubescent inside; ovules (8–)12–20(–24) per ovary; style 3–7 mm, (sometimes pubescent). |
(erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
Seeds | flattened. |
(dark brown), flattened, (2–3 mm). |
2n | = 10. |
= 10, 16, 24. |
Physaria montana |
Physaria acutifolia |
|
Phenology | Flowering Apr–Jun(-Aug). | Flowering May–Jun(-Jul). |
Habitat | Banks, rock outcrops, stony slopes and benchlands, from plains into mountains, in sagebrush, open scrub oak, pinyon-juniper woodland, ponderosa pine, Douglas fir on granitic, often gravelly, non-calcareous soils, rarely on calcareous soils | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities |
Elevation | 1000-3300 m (3300-10800 ft) | 1500-3500 m (4900-11500 ft) |
Distribution |
AZ; CO; NE; NM; SD; WY
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AZ; CO; ID; MT; NM; NV; SD; UT; WY
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Discussion | Physaria montana is a rather variable species that in southwestern Colorado morphologically approaches P. rectipes and in eastern Wyoming approaches P. curvipes; it is unusual in the genus for its frequent presence on igneous, non-calcareous soils. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 650. | FNA vol. 7, p. 624. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Vesicaria montana, Alyssum grayanum, Lesquerella montana, Lesquerella montana var. suffruticosa, Lesquerella rosulata | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa |
Name authority | (A. Gray) Greene: Fl. Francisc., 249. (1891) | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) |
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