Physaria ludoviciana |
Physaria spatulata |
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foothill bladderpod, silver bladderpod |
alpine bladderpod, spatula-leaf bladderpod |
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Habit | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). | Perennials; caudex simple, (relatively small); sparsely to moderately pubescent, trichomes 4- or 5-rayed, rays furcate or bifurcate, not fused, (tuberculate). |
Stems | few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
simple from base, erect to decumbent, (well-exserted beyond basal leaves, loosely spreading), 0.3–1.2 dm. |
Basal leaves | (erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
(erect to prostrate, petiole distinct from blade); blade (inner) spatulate to oblanceolate, or (outer) oblanceolate or orbicular, 1.5–4 cm, margins entire (rarely folded). |
Cauline leaves | blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
blade spatulate, distinctly different from basal. |
Racemes | compact, (elongated and loose in fruit, densely-flowered). |
moderately dense, (6–20-flowered). |
Flowers | sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
sepals (pale yellow), elliptic, 3.5–5 mm; petals lingulate, 6–9 mm. |
Fruiting pedicels | (usually recurved), (5–)10–20(–25) mm. |
(strongly sigmoid), 10–20 mm (2 times longer than fruits). |
Fruits | subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
lanceolate or orbicular, slightly inflated, (2.5–)3–6 mm, (apex usually strongly narrowed); valves pubescent, trichomes sparse and closely appressed to surface; ovules 4–8 per ovary; style 2.5–6 mm (usually ± equal in length to mature fruit). |
Seeds | slightly flattened. |
plump. |
2n | = 10, 20, 30. |
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Physaria ludoviciana |
Physaria spatulata |
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Phenology | Flowering Apr–Jul(-Aug). | Flowering May-early Jul. |
Habitat | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands | Grasslands, subalpine meadows, sagebrush, scattered pines, fellfields, calcareous (sometimes alkaline) substrates |
Elevation | 0-1900 m (0-6200 ft) | 900-2900 m (3000-9500 ft) |
Distribution |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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MT; ND; NE; SD; WY; AB; SK
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Discussion | Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 649. | FNA vol. 7, p. 663. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana | Lesquerella spatulata, Lesquerella alpina var. spatulata, Lesquerella nodosa, P. reediana subsp. spatulata, P. reediana var. spatulata |
Name authority | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | (Rydberg) Grady & O’Kane: Novon 17: 190. (2007) |
Web links |