Physaria ludoviciana |
Physaria pulvinata |
|
---|---|---|
foothill bladderpod, silver bladderpod |
cushion bladderpod |
|
Habit | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). | Perennials; caudex (buried), branched, (dense, forming hard mats); densely pubescent, trichomes (subsessile), 8–13-rayed, rays usually furcate, distinct, (umbonate, usually tuberculate, less so over umbo). |
Stems | few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
several (to several hundred) from base, erect, (each terminating in a tufted cluster of leaves), to 7 dm. |
Basal leaves | (erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
(petiole not differentiated from blade); blade narrowly elliptic to narrowly linear-oblanceolate, (0.8–)1–1.5 cm, (base cuneate), margins entire. |
Cauline leaves | blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
similar to basal, blade sometimes linear, (apex acute). |
Racemes | compact, (elongated and loose in fruit, densely-flowered). |
dense, (often ± subumbellate, somewhat elongated in fruit). |
Flowers | sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
sepals narrowly elliptic, 2.5–3.5(–4) mm, (not keeled); petals narrowly spatulate, 4–7 mm. |
Fruiting pedicels | (usually recurved), (5–)10–20(–25) mm. |
(strongly sigmoid), 5–10 mm. |
Fruits | subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
ellipsoid, compressed, 4–6 mm; valves densely pubescent, trichomes appressed; ovules 2 per ovary; style 2–3.5 mm. |
Seeds | slightly flattened. |
flattened, (oval). |
2n | = 10, 20, 30. |
|
Physaria ludoviciana |
Physaria pulvinata |
|
Phenology | Flowering Apr–Jul(-Aug). | Flowering late May–Jun. |
Habitat | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands | Gray, argillaceous shale outcrops with sagebrush and junipers |
Elevation | 0-1900 m (0-6200 ft) | 2300-2600 m (7500-8500 ft) |
Distribution |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
|
CO |
Discussion | Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria pulvinata is known from an area surrounded by a pygmy forest of Utah juniper in Dolores and San Miguel Counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 649. | FNA vol. 7, p. 659. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana | |
Name authority | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | O’Kane & Reveal: Brittonia 58: 74, fig. 1. (2006) |
Web links |