Physaria ludoviciana |
Physaria pachyphylla |
|
---|---|---|
foothill bladderpod, silver bladderpod |
thick-leaf bladderpod |
|
Habit | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). | Perennials; caudex usually simple, rarely branched, (loosely mounded, rosette-like growth); densely (silvery or gray) pubescent, trichomes (sessile), 5-rayed, rays bifurcate, slightly fused near base of main rays, (tuberculate throughout, less over umbo). |
Stems | few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
several from base, decumbent to prostrate, (well-exserted beyond basal leaves), 0.2–0.5 dm. |
Basal leaves | (erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
(petiole differentiated from blade); blade (slightly cupped, leathery, nearly 1 mm thick), oblanceolate to orbicular, 1.2–2 cm, margins entire, (apex acute). |
Cauline leaves | blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
blade spatulate, similar to basal. |
Racemes | compact, (elongated and loose in fruit, densely-flowered). |
dense, (subumbellate). |
Flowers | sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
sepals (pale yellow), elliptic to oblong, 3.5–4.0 mm, (median pair somewhat thickened apically, cucullate); petals lingulate, 5–6 mm. |
Fruiting pedicels | (usually recurved), (5–)10–20(–25) mm. |
(ascending, curved), 5–7 mm. |
Fruits | subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
globose or ellipsoid, slightly inflated (with slight apical constriction), 3–6 mm; valves pubescent, trichomes closely appressed; ovules 8 per ovary; style 1–3 mm (shorter than mature fruit). |
Seeds | slightly flattened. |
plump, (oblong). |
2n | = 10, 20, 30. |
|
Physaria ludoviciana |
Physaria pachyphylla |
|
Phenology | Flowering Apr–Jul(-Aug). | Flowering Jun–Jul. |
Habitat | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands | Barren areas of mixed white, pink, or reddish limestone and diatomaceous earth |
Elevation | 0-1900 m (0-6200 ft) | 1300-1600 m (4300-5200 ft) |
Distribution |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
|
MT |
Discussion | Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria pachyphylla is known from the Pryor Mountain Desert near the Wyoming state line. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 649. | FNA vol. 7, p. 656. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana | |
Name authority | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | O’Kane & Grady: Novon 17: 187, fig. 4. (2007) |
Web links |