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foothill bladderpod, silver bladderpod

roundleaf bladderpod

Habit Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). Perennials; caudex simple or branched, (thickened by persistent leaf bases); densely pubescent (foliage usually scabrous), trichomes (sessile or short-stalked), several-rayed, rays furcate near base, (usually strongly umbonate, roughly tuberculate, less so over umbo).
Stems

few from base, erect with outer usually decumbent, 1–3.5(–5) dm.

few to several from base, erect or outer decumbent, 0.5–2.5 dm.

Basal leaves

(erect);

blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes).

blade suborbicular to elliptic or ovate or deltate, 0.5–2(–6.5) cm, margins entire or shallowly dentate.

Cauline leaves

blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute.

(proximal shortly petiolate, distal usually sessile);

blade narrowly elliptic or obovate, (0.5–)1–2.5(–4) cm, margins entire.

Racemes

compact, (elongated and loose in fruit, densely-flowered).

compact, (± subumbellate to densely corymbiform, elongated or not).

Flowers

sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate);

petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw).

sepals ± elliptic, 4.5–7(–8.5) mm, (median pair thickened apically);

petals (sometimes white), suborbicular to obovate or obdeltate, 6.5–15 mm, (base narrowing to broad claw, apex sometimes emarginated).

Fruiting pedicels

(usually recurved), (5–)10–20(–25) mm.

(usually spreading at right angles, sometimes nearly erect, ± straight), 5–15(–20) mm, (stout).

Fruits

subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm;

valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile;

ovules (4–)8–12(–16) per ovary;

style 3–4.5(–6.5) mm.

(sessile or shortly stipitate, less than 1 mm), subglobose to broadly ellipsoid, inflated or slightly compressed (terete or subterete), (4–)5–8(–9) mm;

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules 8–16 per ovary;

style 4–8(–9) mm.

Seeds

slightly flattened.

flattened.

2n

= 10, 20, 30.

Physaria ludoviciana

Physaria ovalifolia

Phenology Flowering Apr–Jul(-Aug).
Habitat Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands
Elevation 0-1900 m (0-6200 ft)
Distribution
from FNA
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; KS; NE; NM; OK; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudices branched (well-developed); petals usually yellow, rarely white, 6.5-12(-14) mm, usually 1.5 times or less as long as sepals; racemes usually not elongated (subumbellate).
subsp. ovalifolia
1. Caudices usually simple; petals white, (9-)11-15 mm, often 2 times as long as sepals; racemes usually elongated.
subsp. alba
Source FNA vol. 7, p. 649. FNA vol. 7, p. 655.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. ovalifolia subsp. alba, P. ovalifolia subsp. ovalifolia
Synonyms Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana Lesquerella ovalifolia, Lesquerella engelmannii subsp. ovalifolia
Name authority (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) (Rydberg) O’Kane & Al-Shehbaz: Novon 12: 326. (2002)
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