Physaria ludoviciana |
Physaria multiceps |
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foothill bladderpod, silver bladderpod |
manyhead bladderpod |
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Habit | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). | Perennials; caudex (buried), branched, (not thickened); densely pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays furcate or bifurcate, (umbonate, rough to finely tuberculate throughout). |
Stems | few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
several from base, prostrate, (slender, sparsely pubescent), 0.5–2 dm. |
Basal leaves | (erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
blade obovate to narrowly elliptic, 1.5–6 cm, margins usually entire, rarely shallowly dentate, (surfaces densely pubescent, often silvery). |
Cauline leaves | blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
blade oblanceolate to spatulate, 0.5–1 cm, margins entire, (surfaces often sparsely pubescent). |
Racemes | compact, (elongated and loose in fruit, densely-flowered). |
(narrow), loose, (elongated in fruit). |
Flowers | sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
sepals (greenish brown, sometimes magenta), linear or elliptic, 4.3–6(–7.5) mm, (median pair thickened apically, cucullate); petals (frequently pink or magenta in distal 1/3–1/2), spatulate to oblanceolate, 6–10(–12) mm, (claw undifferentiated from blade). |
Fruiting pedicels | (usually recurved), (5–)10–20(–25) mm. |
(ascending to somewhat spreading, straight to slightly curved), 4–8(–12) mm. |
Fruits | subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
broadly ovoid to suborbicular, inflated, (terete or, often, slightly angustiseptate), 3–6mm; valves sparsely pubescent; ovules usually 4, rarely 6–8 per ovary; style 3–6.5 mm. |
Seeds | slightly flattened. |
plump. |
2n | = 10, 20, 30. |
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Physaria ludoviciana |
Physaria multiceps |
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Phenology | Flowering Apr–Jul(-Aug). | Flowering May–Jul. |
Habitat | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands | Douglas-fir or spruce woodlands, limestone ridges, damp open slopes, soil pockets among rocks, crevices of rocks, decomposed calcareous rocks |
Elevation | 0-1900 m (0-6200 ft) | 2400-2900 m (7900-9500 ft) |
Distribution |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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ID; UT; WY |
Discussion | Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 649. | FNA vol. 7, p. 651. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana | Lesquerella multiceps |
Name authority | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | (Maguire) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
Web links |