Physaria ludoviciana |
Physaria grahamii |
|
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foothill bladderpod, silver bladderpod |
Graham's twinpod |
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Habit | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. |
Stems | few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
Basal leaves | (erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
(outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
Cauline leaves | blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
Racemes | compact, (elongated and loose in fruit, densely-flowered). |
loose, (elongated). |
Flowers | sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
Fruiting pedicels | (usually recurved), (5–)10–20(–25) mm. |
(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
Fruits | subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
Seeds | slightly flattened. |
plump, (suborbicular). |
2n | = 10, 20, 30. |
|
Physaria ludoviciana |
Physaria grahamii |
|
Phenology | Flowering Apr–Jul(-Aug). | Flowering May–Jun. |
Habitat | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay |
Elevation | 0-1900 m (0-6200 ft) | 2100-2900 m (6900-9500 ft) |
Distribution |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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UT |
Discussion | Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 649. | FNA vol. 7, p. 642. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda |
Name authority | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) |
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