Physaria ludoviciana |
Physaria arenosa |
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foothill bladderpod, silver bladderpod |
Great Plains bladderpod |
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Habit | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). | Perennials or, rarely, annuals; caudex simple or branched; ± densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (usually spreading), distinct or slightly fused at base, furcate or bifurcate, (tuberculate). | ||||
Stems | few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
simple or few from base, prostrate or straggling to erect, (sometimes purplish, usually unbranched), (0.5–)1–2(–3) dm. |
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Basal leaves | (erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
blade oblanceolate, 1.5–5(–7) cm, margins entire or shallowly dentate, (flat). |
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Cauline leaves | blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
blade elliptic to linear, (0.5–)1–2.5(–3) cm, margins usually entire. |
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Racemes | compact, (elongated and loose in fruit, densely-flowered). |
(secund), dense, (elongated in fruit). |
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Flowers | sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
sepals elliptic or oblong, 4–6(–7) mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (often red or lavender when dried), obovate, 6–8.5(–9.5) mm, (narrowing to broad claw). |
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Fruiting pedicels | (usually recurved), (5–)10–20(–25) mm. |
(usually sharply recurved, sometimes divaricate-spreading or nearly horizontal), 5–15(–20) mm, (stout). |
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Fruits | subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
subglobose, obovoid, or broadly ellipsoid, slightly inflated, (3.5–)4–5.5(–6.5) mm; valves densely pubescent outside, trichomes spreading or closely appressed, rarely sparsely pubescent inside; ovules (4–)8(–10) per ovary; style (slender), 3–5.5(–6.5) mm. |
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Seeds | slightly flattened. |
slightly flattened. |
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2n | = 10, 20, 30. |
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Physaria ludoviciana |
Physaria arenosa |
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Phenology | Flowering Apr–Jul(-Aug). | |||||
Habitat | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands | |||||
Elevation | 0-1900 m (0-6200 ft) | |||||
Distribution |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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CO; MT; ND; NE; SD; WY; AB; MB; SK
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Discussion | Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 649. | FNA vol. 7, p. 626. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana | Vesicaria arenosa, Lesquerella arenosa, Lesquerella argentea var. arenosa, Lesquerella ludoviciana var. arenosa | ||||
Name authority | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | (Richardson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | ||||
Web links |