Physaria ludoviciana |
Physaria alpina |
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foothill bladderpod, silver bladderpod |
Avery Peak or alpine twinpod, Avery Peak twinpod |
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Habit | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays usually furcate, sometimes bifurcate, (rough-tuberculate throughout). | Perennials; (with a long taproot), caudex usually buried, simple, (enlarged, covered with marcescent leaf bases, crown rosulate and horizontal to somewhat ascending, forming a dense crown at apex of caudex); (silvery) pubescent throughout, trichomes (sessile or stipitate), 5–8-rayed, rays furcate or bifurcate, (rounded to umbonate, strongly tuberculate, less so or smooth over center). |
Stems | few from base, erect with outer usually decumbent, 1–3.5(–5) dm. |
few from base, decumbent, (arising laterally proximal to current season’s leaves), 0.3–0.8 dm. |
Basal leaves | (erect); blade narrowly lanceolate to linear or (outer) oblanceolate, (1–)2–6(–9) cm, margins usually entire, rarely shallowly dentate, (inner involute, outer usually flat, base usually with some simple trichomes). |
(petiole slender); blade broadly obovate, or deltate to ovate or narrower, 1.5–3.5 cm, (base abruptly to gradually narrowed to petiole), margins entire or obscurely few-toothed, (apex usually obtuse, nearly acute in narrower leaves). |
Cauline leaves | blade narrowly oblanceolate to linear, (1–)2–4(–8) cm, margins flat or involute. |
(2–5 per stem); blade oblanceolate to spatulate, similar to basal, margins entire, (apex acute). |
Racemes | compact, (elongated and loose in fruit, densely-flowered). |
loose, (3–6-flowered). |
Flowers | sepals oblong to broadly elliptic, 4–7(–8) mm, (lateral pair subsaccate, median pair cucullate); petals oblanceolate or obovate, (5–)6.5–9.5(–11) mm, (claw undifferentiated from blade, or blade gradually narrowed to claw). |
sepals narrowly oblong to linear, 7–9 mm; petals (erect), spatulate, 10–12(–15) mm. |
Fruiting pedicels | (usually recurved), (5–)10–20(–25) mm. |
(widely spreading to ascending, slightly curved or straight), 7–11 mm. |
Fruits | subglobose or obovoid, usually inflated, sometimes slightly compressed, (3–)4–6 mm; valves densely pubescent, trichomes spreading, usually pubescent inside, trichomes appressed, sessile; ovules (4–)8–12(–16) per ovary; style 3–4.5(–6.5) mm. |
(usually purplish in age), didymous, irregular and somewhat angular, not highly inflated, 4–11 × 10–13 mm, (coriaceous, papery, shallowly grooved distally and on sides, tapered and narrowed toward replum, base obtuse to truncate, apex with broad sinus to nearly truncate); valves (retaining seeds after dehiscence), densely pubescent, not silvery; replum elliptic to obovate, as wide as or wider than fruit, base rounded, margins sparsely pubescent or glabrous, apex rounded (with funicles); ovules 4 per ovary; style 5–7 mm, (glabrous). |
Seeds | slightly flattened. |
flattened. |
2n | = 10, 20, 30. |
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Physaria ludoviciana |
Physaria alpina |
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Phenology | Flowering Apr–Jul(-Aug). | Flowering Jun–Jul. |
Habitat | Sandy or gravelly soils, steep hillsides, prairie pastures, clay slopes, limestone outcrops, sand dunes, open plains, sandy bluffs, rocky flats, white tuff sands | Whitish or red substrates from limestone or dolomite, ridge crests, rocky alpine tundra and open areas |
Elevation | 0-1900 m (0-6200 ft) | 3500-4000 m (11500-13100 ft) |
Distribution |
AZ; CA; CO; IA; IL; KS; MN; MT; ND; NE; NM; NV; OK; SD; UT; WI; WY
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CO
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Discussion | Material previously reported as Physaria ludoviciana from Canada (Alberta, Manitoba, Saskatchewan) is here included in 6a. P. arenosa subsp. arenosa. Lesquerella argentea (Pursh) MacMillan is a later homonym that has been used for P. ludoviciana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 649. | FNA vol. 7, p. 624. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | Alyssum ludovicianum, Lesquerella ludoviciana, Vesicaria ludoviciana | |
Name authority | (Nuttall) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | Rollins: Brittonia 33: 339. (1981) |
Web links |