Physaria kingii |
Physaria pallida |
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King bladderpod, King's bladder-pod |
white bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (not thickened); usually densely pubescent, trichomes (sessile or short-stalked), 3–7-rayed, rays distinct or slightly fused at base, typically furcate near base, bifurcate or 3-partite, (not to slightly umbonate, smooth or moderately to strongly tuberculate). | Annuals (winter); with a fine taproot; sparsely pubescent, trichomes (minute), 3- or 4-rayed, rays furcate or, sometimes, trifurcate. | ||||||||||||||||||||||||
Stems | few to several from base, prostrate to decumbent or erect, 0.5–2(–4) dm. |
few to several from base, erect, (from basal leaf cluster, branched distally, flowering branches slender, subtended by bracts), 3–6 dm. |
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Basal leaves | blade suborbicular to narrowly or broadly oblanceolate to broadly elliptic or rhombic, (1.2–)2–6(–8) cm, (base usually abruptly narrowed to petiole), margins entire, sinuate, or lobed. |
blade oblanceolate or broadly obovate, to 10 cm, margins usually sinuate-dentate or entire, sometimes lobed. |
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Cauline leaves | (proximal petiolate, distal sessile); blade obovate or elliptic to spatulate, 0.5–2 cm, margins entire. |
(proximal shortly petiolate, distal sessile); blade oblanceolate to narrowly oblong, similar to basal, (distal with base slightly cuneate). |
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Racemes | (usually not secund), dense, (sometimes elongated in fruit). |
paniculate, (rachises and pedicels more densely pubescent than proximal leaves). |
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Flowers | sepals lanceolate, 4–6(–7) mm; petals (yellow, cream-yellow, cream-white, or white), obovate to oblanceolate, 6–13 mm, (claw weakly differentiated from blade). |
sepals elliptic, 3–7 mm, (median pair slightly thickened apically, cucullate); petals (white), broadly ovate, to 12 mm, (narrowing gradually to short claw). |
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Fruiting pedicels | (erect to divaricate-ascending or recurved, erect in distal 1/3, usually sigmoid, sometimes straight or slightly curved), 4.5–10(–15) mm. |
(widely divaricate-ascending and straight, or slightly recurved), 10–15 mm, (slender, pubescent). |
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Fruits | (sessile or substipitate), subglobose, obovoid, or ellipsoid, compressed (sometimes slightly angustiseptate), 3–9 mm, (rigid, apex truncate, retuse, or rounded-acute); valves sparsely or densely pubescent, sometimes sparsely pubescent inside; (septum sometimes fenestrate, perforate, or obsolete); ovules 4–16 per ovary; style 1–9 mm. |
(widely spreading to nearly pendent in age, shortly stipitate), globose or subglobose, not or slightly inflated, 3–10 mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–12 per ovary; style ca. 2 mm, (slender, fragile). |
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Seeds | flattened (sometimes slightly). |
flattened. |
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2n | = 12. |
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Physaria kingii |
Physaria pallida |
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Phenology | Flowering Apr–May. | |||||||||||||||||||||||||
Habitat | Grassy openings of small glade prairies, outcrops | |||||||||||||||||||||||||
Elevation | 90 m (300 ft) | |||||||||||||||||||||||||
Distribution |
AZ; CA; ID; NV; OR; UT; nw Mexico
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TX |
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Discussion | Subspecies 7 (7 in the flora). The Physaria kingii complex is in need of further study. It is widespread in the western United States, mostly in montane environments. This treatment recognizes a highly variable species with generally well-marked, geographically coherent subspecies. Hybridization may be involved in some of the subspecies, especially in subsp. kaibabensis, where molecular data indicate intra-individual genetic variation (pers. obs.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria pallida is known from the Weches Formation in San Augustine County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 645. | FNA vol. 7, p. 656. | ||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||
Synonyms | Vesicaria kingii, Lesquerella kingii | Vesicaria grandiflora var. pallida, Alyssum pallidum, Lesquerella pallida, Vesicaria pallida | ||||||||||||||||||||||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | (Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | ||||||||||||||||||||||||
Web links |