Physaria kingii |
Physaria multiceps |
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King bladderpod, King's bladder-pod |
manyhead bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (not thickened); usually densely pubescent, trichomes (sessile or short-stalked), 3–7-rayed, rays distinct or slightly fused at base, typically furcate near base, bifurcate or 3-partite, (not to slightly umbonate, smooth or moderately to strongly tuberculate). | Perennials; caudex (buried), branched, (not thickened); densely pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays furcate or bifurcate, (umbonate, rough to finely tuberculate throughout). | ||||||||||||||||||||||||
Stems | few to several from base, prostrate to decumbent or erect, 0.5–2(–4) dm. |
several from base, prostrate, (slender, sparsely pubescent), 0.5–2 dm. |
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Basal leaves | blade suborbicular to narrowly or broadly oblanceolate to broadly elliptic or rhombic, (1.2–)2–6(–8) cm, (base usually abruptly narrowed to petiole), margins entire, sinuate, or lobed. |
blade obovate to narrowly elliptic, 1.5–6 cm, margins usually entire, rarely shallowly dentate, (surfaces densely pubescent, often silvery). |
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Cauline leaves | (proximal petiolate, distal sessile); blade obovate or elliptic to spatulate, 0.5–2 cm, margins entire. |
blade oblanceolate to spatulate, 0.5–1 cm, margins entire, (surfaces often sparsely pubescent). |
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Racemes | (usually not secund), dense, (sometimes elongated in fruit). |
(narrow), loose, (elongated in fruit). |
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Flowers | sepals lanceolate, 4–6(–7) mm; petals (yellow, cream-yellow, cream-white, or white), obovate to oblanceolate, 6–13 mm, (claw weakly differentiated from blade). |
sepals (greenish brown, sometimes magenta), linear or elliptic, 4.3–6(–7.5) mm, (median pair thickened apically, cucullate); petals (frequently pink or magenta in distal 1/3–1/2), spatulate to oblanceolate, 6–10(–12) mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (erect to divaricate-ascending or recurved, erect in distal 1/3, usually sigmoid, sometimes straight or slightly curved), 4.5–10(–15) mm. |
(ascending to somewhat spreading, straight to slightly curved), 4–8(–12) mm. |
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Fruits | (sessile or substipitate), subglobose, obovoid, or ellipsoid, compressed (sometimes slightly angustiseptate), 3–9 mm, (rigid, apex truncate, retuse, or rounded-acute); valves sparsely or densely pubescent, sometimes sparsely pubescent inside; (septum sometimes fenestrate, perforate, or obsolete); ovules 4–16 per ovary; style 1–9 mm. |
broadly ovoid to suborbicular, inflated, (terete or, often, slightly angustiseptate), 3–6mm; valves sparsely pubescent; ovules usually 4, rarely 6–8 per ovary; style 3–6.5 mm. |
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Seeds | flattened (sometimes slightly). |
plump. |
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Physaria kingii |
Physaria multiceps |
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Phenology | Flowering May–Jul. | |||||||||||||||||||||||||
Habitat | Douglas-fir or spruce woodlands, limestone ridges, damp open slopes, soil pockets among rocks, crevices of rocks, decomposed calcareous rocks | |||||||||||||||||||||||||
Elevation | 2400-2900 m (7900-9500 ft) | |||||||||||||||||||||||||
Distribution |
AZ; CA; ID; NV; OR; UT; nw Mexico
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ID; UT; WY |
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Discussion | Subspecies 7 (7 in the flora). The Physaria kingii complex is in need of further study. It is widespread in the western United States, mostly in montane environments. This treatment recognizes a highly variable species with generally well-marked, geographically coherent subspecies. Hybridization may be involved in some of the subspecies, especially in subsp. kaibabensis, where molecular data indicate intra-individual genetic variation (pers. obs.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 645. | FNA vol. 7, p. 651. | ||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||
Synonyms | Vesicaria kingii, Lesquerella kingii | Lesquerella multiceps | ||||||||||||||||||||||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | (Maguire) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | ||||||||||||||||||||||||
Web links |