Physaria kingii |
Physaria fendleri |
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King bladderpod, King's bladder-pod |
Fendler bladderpod, Fendler's bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (not thickened); usually densely pubescent, trichomes (sessile or short-stalked), 3–7-rayed, rays distinct or slightly fused at base, typically furcate near base, bifurcate or 3-partite, (not to slightly umbonate, smooth or moderately to strongly tuberculate). | Perennials; caudex branched, (sometimes woody at base); densely (silvery) pubescent, trichomes (sessile or short-stalked), several-rayed, rays not furcate, fused (webbed) ca. 1/2 their length, (tuberculate throughout or tubercles scarce or absent over center). | ||||||||||||||||||||||||
Stems | few to several from base, prostrate to decumbent or erect, 0.5–2(–4) dm. |
several from base, erect or laterally decumbent, (usually unbranched), (0.3–)0.5–2.5(–4) dm. |
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Basal leaves | blade suborbicular to narrowly or broadly oblanceolate to broadly elliptic or rhombic, (1.2–)2–6(–8) cm, (base usually abruptly narrowed to petiole), margins entire, sinuate, or lobed. |
blade linear to somewhat elliptic, 1–4(–8) cm, margins entire or coarsely dentate. |
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Cauline leaves | (proximal petiolate, distal sessile); blade obovate or elliptic to spatulate, 0.5–2 cm, margins entire. |
(shortly petiolate); blade usually linear to narrowly oblanceolate, rarely elliptic to rhombic, 0.5–2.5 cm, (base narrowing to petiole), margins entire or remotely dentate (sometimes involute). |
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Racemes | (usually not secund), dense, (sometimes elongated in fruit). |
loose to somewhat dense. |
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Flowers | sepals lanceolate, 4–6(–7) mm; petals (yellow, cream-yellow, cream-white, or white), obovate to oblanceolate, 6–13 mm, (claw weakly differentiated from blade). |
sepals elliptic to oblong, 5–8 mm, (lateral pair not saccate, median pair often thickened apically, ± cucullate); petals (usually orange or orange-yellow at junction of blade and claw, sometimes also with orange guidelines), obdeltate to obovate, 8–12 mm, (claw relatively short). |
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Fruiting pedicels | (erect to divaricate-ascending or recurved, erect in distal 1/3, usually sigmoid, sometimes straight or slightly curved), 4.5–10(–15) mm. |
(divaricate-spreading to erect, usually straight or slightly curved, occasionally sigmoid), 8–20(–40) mm. |
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Fruits | (sessile or substipitate), subglobose, obovoid, or ellipsoid, compressed (sometimes slightly angustiseptate), 3–9 mm, (rigid, apex truncate, retuse, or rounded-acute); valves sparsely or densely pubescent, sometimes sparsely pubescent inside; (septum sometimes fenestrate, perforate, or obsolete); ovules 4–16 per ovary; style 1–9 mm. |
globose, broadly ellipsoid, or ovoid, not or slightly inflated, 5–8 mm, (firm, apex usually acute); valves (not retaining seeds after dehiscence, often reddish in age), glabrous throughout; replum as wide as or wider than fruit; ovules (12–)20–32(–40) per ovary; style (2–)3–6 mm. |
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Seeds | flattened (sometimes slightly). |
flattened. |
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2n | = 12, 14, 24. |
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Physaria kingii |
Physaria fendleri |
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Phenology | Flowering Mar–May. | |||||||||||||||||||||||||
Habitat | Limestone outcrops, gypseous hills, gravels, sandy washes, rocky slopes, bluffs, shallow drainage areas, plains and desert shrub areas | |||||||||||||||||||||||||
Elevation | 100-2000 m (300-6600 ft) | |||||||||||||||||||||||||
Distribution |
AZ; CA; ID; NV; OR; UT; nw Mexico
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AZ; CO; KS; NM; OK; TX; UT; Mexico (Chihuahua, Coahuila, Nuevo León)
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Discussion | Subspecies 7 (7 in the flora). The Physaria kingii complex is in need of further study. It is widespread in the western United States, mostly in montane environments. This treatment recognizes a highly variable species with generally well-marked, geographically coherent subspecies. Hybridization may be involved in some of the subspecies, especially in subsp. kaibabensis, where molecular data indicate intra-individual genetic variation (pers. obs.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In dry areas, Physaria fendleri may flower following suitable rains at any time of the year. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 645. | FNA vol. 7, p. 636. | ||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||
Synonyms | Vesicaria kingii, Lesquerella kingii | Vesicaria fendleri, Alyssum fendleri, Alyssum stenophyllum, Lesquerella fendleri, Lesquerella foliacea, Lesquerella praecox, Lesquerella stenophylla, Vesicaria stenophylla, Vesicaria stenophylla var. diffusa, Vesicaria stenophylla var. humilis, Vesicaria stenophylla var. procera | ||||||||||||||||||||||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||||||||||||||||||||||
Web links |