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King bladderpod, King's bladder-pod

tuft twinpod

Habit Perennials; caudex usually simple, sometimes branched, (not thickened); usually densely pubescent, trichomes (sessile or short-stalked), 3–7-rayed, rays distinct or slightly fused at base, typically furcate near base, bifurcate or 3-partite, (not to slightly umbonate, smooth or moderately to strongly tuberculate). Perennials; caudex usually simple, rarely branched, (enlarged with persistent leaf bases, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays typically furcate, (fused at base, arms slender, tuberculate throughout).
Stems

few to several from base, prostrate to decumbent or erect, 0.5–2(–4) dm.

several from base, decumbent to ascending, (arising laterally beneath a dense rosette), less than 0.1 dm.

Basal leaves

blade suborbicular to narrowly or broadly oblanceolate to broadly elliptic or rhombic, (1.2–)2–6(–8) cm, (base usually abruptly narrowed to petiole), margins entire, sinuate, or lobed.

(petiole slender);

blade (horizontal on the ground), obovate, 0.5–1.5 cm (width 4–8 mm, base tapering abruptly to petiole), margins entire, (apex usually acute, surfaces silvery from a dense incrustation of appressed, stellate trichomes).

Cauline leaves

(proximal petiolate, distal sessile);

blade obovate or elliptic to spatulate, 0.5–2 cm, margins entire.

blade oblanceolate, 0.5–1 cm (width 2–3 mm), margins entire, (surfaces densely stellate pubescent).

Racemes

(usually not secund), dense, (sometimes elongated in fruit).

congested, (subumbellate, often almost sessile, barely exceeding basal leaves).

Flowers

sepals lanceolate, 4–6(–7) mm;

petals (yellow, cream-yellow, cream-white, or white), obovate to oblanceolate, 6–13 mm, (claw weakly differentiated from blade).

sepals (yellowish green), narrowly lanceolate, 4–5 mm;

petals (erect), oblanceolate, 6–7 mm, (claw weakly differentiated from blade).

Fruiting pedicels

(erect to divaricate-ascending or recurved, erect in distal 1/3, usually sigmoid, sometimes straight or slightly curved), 4.5–10(–15) mm.

(divaricate, straight), 5–10 mm.

Fruits

(sessile or substipitate), subglobose, obovoid, or ellipsoid, compressed (sometimes slightly angustiseptate), 3–9 mm, (rigid, apex truncate, retuse, or rounded-acute);

valves sparsely or densely pubescent, sometimes sparsely pubescent inside; (septum sometimes fenestrate, perforate, or obsolete);

ovules 4–16 per ovary;

style 1–9 mm.

didymous, ovate, inflated, 4.8–6 × 6–10 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), pubescent, trichomes loosely spreading;

replum obovate, 3–4 mm, as wide as or wider than fruit;

ovules 8 per ovary (2–4 abortive);

style 4–6 mm.

Seeds

flattened (sometimes slightly).

flattened.

Physaria kingii

Physaria condensata

Phenology Flowering May–Jun.
Habitat Calcareous knolls and ridges, clay banks, limey slopes, shaley hills, clay patches
Elevation 1800-2400 m (5900-7900 ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Subspecies 7 (7 in the flora).

The Physaria kingii complex is in need of further study. It is widespread in the western United States, mostly in montane environments. This treatment recognizes a highly variable species with generally well-marked, geographically coherent subspecies. Hybridization may be involved in some of the subspecies, especially in subsp. kaibabensis, where molecular data indicate intra-individual genetic variation (pers. obs.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits slightly wider than long, apices truncate or retuse, valves pubescent inside
→ 2
1. Fruits as wide as or longer than wide, apices rounded-acute, valves glabrous inside
→ 3
2. Basal leaf blades: margins ± entire (sometimes slightly lobed or widened at base); California, Idaho, Nevada, Oregon.
subsp. kingii
2. Basal leaf blades: margins sinuate or lobed, or, sometimes, lyrate; Wallowa and Elkhorn mountains, Oregon.
subsp. diversifolia
3. Fruiting pedicels recurved.
subsp. cobrensis
3. Fruiting pedicels not recurved (divaricate-ascending or ± erect, straight or sigmoid)
→ 4
4. Petals cream-white or white; styles 1-2 mm; Kaibab Plateau, n Arizona.
subsp. kaibabensis
4. Petals yellow (occasionally cream-yellow or cream-white on Kaibab Plateau, Arizona); styles (4-)4.5-9 mm; n Arizona (including Kaibab Plateau), California, Nevada, Utah
→ 5
5. Plants erect; styles 6-9 mm; ovules 4-8 per ovary; se California.
subsp. bernardina
5. Plants ascending, erect, decumbent, or prostrate; styles (4-)4.5-7 mm; ovules (6-)8-16 per ovary; n Arizona, e California, s Nevada, Utah
→ 6
6. Plants prostrate, decumbent, or erect; racemes not or somewhat secund in fruit; ovules usually 8-16 per ovary; n Arizona, e California, s Nevada, Utah.
subsp. latifolia
6. Plants ascending; racemes secund in fruit; ovules (6-)8-12 per ovary; n Utah.
subsp. utahensis
Source FNA vol. 7, p. 645. FNA vol. 7, p. 631.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. kingii subsp. bernardina, P. kingii subsp. cobrensis, P. kingii subsp. diversifolia, P. kingii subsp. kaibabensis, P. kingii subsp. kingii, P. kingii subsp. latifolia, P. kingii subsp. utahensis
Synonyms Vesicaria kingii, Lesquerella kingii
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) Rollins: Rhodora 41: 407, plate 556, figs. 1, 9, 10. (1939)
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