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King bladderpod, King's bladder-pod

arctic bladderpod

Habit Perennials; caudex usually simple, sometimes branched, (not thickened); usually densely pubescent, trichomes (sessile or short-stalked), 3–7-rayed, rays distinct or slightly fused at base, typically furcate near base, bifurcate or 3-partite, (not to slightly umbonate, smooth or moderately to strongly tuberculate). Perennials; caudex simple or branched, (woody, cespitose); ± densely pubescent, trichomes (sessile or short-stalked), several-rayed, rays distinct, furcate or bifurcate, (somewhat umbonate, finely tuberculate to ± smooth).
Stems

few to several from base, prostrate to decumbent or erect, 0.5–2(–4) dm.

simple or few to several from base, erect to spreading or prostrate, 0.5–2(–3) dm.

Basal leaves

blade suborbicular to narrowly or broadly oblanceolate to broadly elliptic or rhombic, (1.2–)2–6(–8) cm, (base usually abruptly narrowed to petiole), margins entire, sinuate, or lobed.

(usually ± rosulate);

blade obovate to oblanceolate, (1–)2–6(–15) cm, margins entire.

Cauline leaves

(proximal petiolate, distal sessile);

blade obovate or elliptic to spatulate, 0.5–2 cm, margins entire.

(sessile or shortly petiolate);

blade oblanceolate or lingulate, 0.5–1.5(–3) cm, margins entire.

Racemes

(usually not secund), dense, (sometimes elongated in fruit).

loose.

Flowers

sepals lanceolate, 4–6(–7) mm;

petals (yellow, cream-yellow, cream-white, or white), obovate to oblanceolate, 6–13 mm, (claw weakly differentiated from blade).

sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate);

petals spatulate, 5–6(–7) mm, (blade gradually narrowed to claw).

Fruiting pedicels

(erect to divaricate-ascending or recurved, erect in distal 1/3, usually sigmoid, sometimes straight or slightly curved), 4.5–10(–15) mm.

(erect to divaricate or ascending), (5–)10–20(–40) mm, (stout).

Fruits

(sessile or substipitate), subglobose, obovoid, or ellipsoid, compressed (sometimes slightly angustiseptate), 3–9 mm, (rigid, apex truncate, retuse, or rounded-acute);

valves sparsely or densely pubescent, sometimes sparsely pubescent inside; (septum sometimes fenestrate, perforate, or obsolete);

ovules 4–16 per ovary;

style 1–9 mm.

subglobose to ellipsoid, uncompressed, 4–6(–9) mm;

valves (not retaining seeds after dehiscence), glabrous or sparsely pubescent outside, trichomes sessile;

replum as wide as or wider than fruit;

ovules (8–)10–14(–16) per ovary;

style 1–2.5(–4) mm.

Seeds

flattened (sometimes slightly).

plump.

2n

= 60.

Physaria kingii

Physaria arctica

Phenology Flowering May–Aug.
Habitat Typically on sand and gravel from calcareous bedrock, river bars and terraces, cliff ledges, scree and talus slopes, often growing after disturbance
Elevation 0-1800 m (0-5900 ft)
Distribution
from FNA
AZ; CA; ID; NV; OR; UT; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AB; BC; MB; NL; NT; NU; QC; YT; Greenland; circumarctic (except n Europe, ne Russia)
[BONAP county map]
Discussion

Subspecies 7 (7 in the flora).

The Physaria kingii complex is in need of further study. It is widespread in the western United States, mostly in montane environments. This treatment recognizes a highly variable species with generally well-marked, geographically coherent subspecies. Hybridization may be involved in some of the subspecies, especially in subsp. kaibabensis, where molecular data indicate intra-individual genetic variation (pers. obs.).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits slightly wider than long, apices truncate or retuse, valves pubescent inside
→ 2
1. Fruits as wide as or longer than wide, apices rounded-acute, valves glabrous inside
→ 3
2. Basal leaf blades: margins ± entire (sometimes slightly lobed or widened at base); California, Idaho, Nevada, Oregon.
subsp. kingii
2. Basal leaf blades: margins sinuate or lobed, or, sometimes, lyrate; Wallowa and Elkhorn mountains, Oregon.
subsp. diversifolia
3. Fruiting pedicels recurved.
subsp. cobrensis
3. Fruiting pedicels not recurved (divaricate-ascending or ± erect, straight or sigmoid)
→ 4
4. Petals cream-white or white; styles 1-2 mm; Kaibab Plateau, n Arizona.
subsp. kaibabensis
4. Petals yellow (occasionally cream-yellow or cream-white on Kaibab Plateau, Arizona); styles (4-)4.5-9 mm; n Arizona (including Kaibab Plateau), California, Nevada, Utah
→ 5
5. Plants erect; styles 6-9 mm; ovules 4-8 per ovary; se California.
subsp. bernardina
5. Plants ascending, erect, decumbent, or prostrate; styles (4-)4.5-7 mm; ovules (6-)8-16 per ovary; n Arizona, e California, s Nevada, Utah
→ 6
6. Plants prostrate, decumbent, or erect; racemes not or somewhat secund in fruit; ovules usually 8-16 per ovary; n Arizona, e California, s Nevada, Utah.
subsp. latifolia
6. Plants ascending; racemes secund in fruit; ovules (6-)8-12 per ovary; n Utah.
subsp. utahensis
Source FNA vol. 7, p. 645. FNA vol. 7, p. 626.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. kingii subsp. bernardina, P. kingii subsp. cobrensis, P. kingii subsp. diversifolia, P. kingii subsp. kaibabensis, P. kingii subsp. kingii, P. kingii subsp. latifolia, P. kingii subsp. utahensis
Synonyms Vesicaria kingii, Lesquerella kingii Alyssum arcticum, Lesquerella arctica, Lesquerella arctica subsp. purshii, Vesicaria arctica, Vesicaria leiocarpa
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) (Wormskjöld ex Hornemann) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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