Physaria kingii |
Physaria angustifolia |
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King bladderpod, King's bladder-pod |
threadleaf bladderpod |
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Habit | Perennials; caudex usually simple, sometimes branched, (not thickened); usually densely pubescent, trichomes (sessile or short-stalked), 3–7-rayed, rays distinct or slightly fused at base, typically furcate near base, bifurcate or 3-partite, (not to slightly umbonate, smooth or moderately to strongly tuberculate). | Annuals; with a fine taproot; ± densely pubescent, trichomes several-rayed, rays distinct or fused at base, bifurcate, (prominently tuberculate throughout). | ||||||||||||||||||||||||
Stems | few to several from base, prostrate to decumbent or erect, 0.5–2(–4) dm. |
simple or few to several from base, erect, (sometimes branched), to 4 dm. |
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Basal leaves | blade suborbicular to narrowly or broadly oblanceolate to broadly elliptic or rhombic, (1.2–)2–6(–8) cm, (base usually abruptly narrowed to petiole), margins entire, sinuate, or lobed. |
blade elliptic to rhombic, 3–8 cm, (base narrowing gradually to petiole), margins entire, repand, coarsely toothed, or pinnatifid. |
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Cauline leaves | (proximal petiolate, distal sessile); blade obovate or elliptic to spatulate, 0.5–2 cm, margins entire. |
(proximal often shortly petiolate, distal sessile); blade linear or narrowly obovate, 1.5–6(–10) cm, margins entire, repand, or shallowly toothed. |
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Racemes | (usually not secund), dense, (sometimes elongated in fruit). |
usually loose. |
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Flowers | sepals lanceolate, 4–6(–7) mm; petals (yellow, cream-yellow, cream-white, or white), obovate to oblanceolate, 6–13 mm, (claw weakly differentiated from blade). |
sepals elliptic or ovate, 4–6 mm, (lateral pair usually subsaccate); petals obovate to obdeltate, 6–10 mm, (apex often emarginate). |
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Fruiting pedicels | (erect to divaricate-ascending or recurved, erect in distal 1/3, usually sigmoid, sometimes straight or slightly curved), 4.5–10(–15) mm. |
(usually divaricate, sometimes horizontal, straight or slightly curved), 8–20 mm. |
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Fruits | (sessile or substipitate), subglobose, obovoid, or ellipsoid, compressed (sometimes slightly angustiseptate), 3–9 mm, (rigid, apex truncate, retuse, or rounded-acute); valves sparsely or densely pubescent, sometimes sparsely pubescent inside; (septum sometimes fenestrate, perforate, or obsolete); ovules 4–16 per ovary; style 1–9 mm. |
not didymous, ± globose, slightly inflated, 4–6 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4 per ovary; style 2–3.5 mm; (stigma expanded). |
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Seeds | flattened (sometimes slightly). |
flattened, (margined). |
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2n | = 10. |
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Physaria kingii |
Physaria angustifolia |
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Phenology | Flowering Apr(-May). | |||||||||||||||||||||||||
Habitat | Shallow limestone-derived soils, sometimes spreading to disturbed sites | |||||||||||||||||||||||||
Elevation | 90-300 m (300-1000 ft) | |||||||||||||||||||||||||
Distribution |
AZ; CA; ID; NV; OR; UT; nw Mexico
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OK; TX |
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Discussion | Subspecies 7 (7 in the flora). The Physaria kingii complex is in need of further study. It is widespread in the western United States, mostly in montane environments. This treatment recognizes a highly variable species with generally well-marked, geographically coherent subspecies. Hybridization may be involved in some of the subspecies, especially in subsp. kaibabensis, where molecular data indicate intra-individual genetic variation (pers. obs.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 645. | FNA vol. 7, p. 625. | ||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||
Synonyms | Vesicaria kingii, Lesquerella kingii | Vesicaria angustifolia, Lesquerella angustifolia, Lesquerella longifolia | ||||||||||||||||||||||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | (Nuttall ex Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | ||||||||||||||||||||||||
Web links |