Physaria kingii |
Physaria alpina |
|||||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
King bladderpod, King's bladder-pod |
Avery Peak or alpine twinpod, Avery Peak twinpod |
|||||||||||||||||||||||||
Habit | Perennials; caudex usually simple, sometimes branched, (not thickened); usually densely pubescent, trichomes (sessile or short-stalked), 3–7-rayed, rays distinct or slightly fused at base, typically furcate near base, bifurcate or 3-partite, (not to slightly umbonate, smooth or moderately to strongly tuberculate). | Perennials; (with a long taproot), caudex usually buried, simple, (enlarged, covered with marcescent leaf bases, crown rosulate and horizontal to somewhat ascending, forming a dense crown at apex of caudex); (silvery) pubescent throughout, trichomes (sessile or stipitate), 5–8-rayed, rays furcate or bifurcate, (rounded to umbonate, strongly tuberculate, less so or smooth over center). | ||||||||||||||||||||||||
Stems | few to several from base, prostrate to decumbent or erect, 0.5–2(–4) dm. |
few from base, decumbent, (arising laterally proximal to current season’s leaves), 0.3–0.8 dm. |
||||||||||||||||||||||||
Basal leaves | blade suborbicular to narrowly or broadly oblanceolate to broadly elliptic or rhombic, (1.2–)2–6(–8) cm, (base usually abruptly narrowed to petiole), margins entire, sinuate, or lobed. |
(petiole slender); blade broadly obovate, or deltate to ovate or narrower, 1.5–3.5 cm, (base abruptly to gradually narrowed to petiole), margins entire or obscurely few-toothed, (apex usually obtuse, nearly acute in narrower leaves). |
||||||||||||||||||||||||
Cauline leaves | (proximal petiolate, distal sessile); blade obovate or elliptic to spatulate, 0.5–2 cm, margins entire. |
(2–5 per stem); blade oblanceolate to spatulate, similar to basal, margins entire, (apex acute). |
||||||||||||||||||||||||
Racemes | (usually not secund), dense, (sometimes elongated in fruit). |
loose, (3–6-flowered). |
||||||||||||||||||||||||
Flowers | sepals lanceolate, 4–6(–7) mm; petals (yellow, cream-yellow, cream-white, or white), obovate to oblanceolate, 6–13 mm, (claw weakly differentiated from blade). |
sepals narrowly oblong to linear, 7–9 mm; petals (erect), spatulate, 10–12(–15) mm. |
||||||||||||||||||||||||
Fruiting pedicels | (erect to divaricate-ascending or recurved, erect in distal 1/3, usually sigmoid, sometimes straight or slightly curved), 4.5–10(–15) mm. |
(widely spreading to ascending, slightly curved or straight), 7–11 mm. |
||||||||||||||||||||||||
Fruits | (sessile or substipitate), subglobose, obovoid, or ellipsoid, compressed (sometimes slightly angustiseptate), 3–9 mm, (rigid, apex truncate, retuse, or rounded-acute); valves sparsely or densely pubescent, sometimes sparsely pubescent inside; (septum sometimes fenestrate, perforate, or obsolete); ovules 4–16 per ovary; style 1–9 mm. |
(usually purplish in age), didymous, irregular and somewhat angular, not highly inflated, 4–11 × 10–13 mm, (coriaceous, papery, shallowly grooved distally and on sides, tapered and narrowed toward replum, base obtuse to truncate, apex with broad sinus to nearly truncate); valves (retaining seeds after dehiscence), densely pubescent, not silvery; replum elliptic to obovate, as wide as or wider than fruit, base rounded, margins sparsely pubescent or glabrous, apex rounded (with funicles); ovules 4 per ovary; style 5–7 mm, (glabrous). |
||||||||||||||||||||||||
Seeds | flattened (sometimes slightly). |
flattened. |
||||||||||||||||||||||||
Physaria kingii |
Physaria alpina |
|||||||||||||||||||||||||
Phenology | Flowering Jun–Jul. | |||||||||||||||||||||||||
Habitat | Whitish or red substrates from limestone or dolomite, ridge crests, rocky alpine tundra and open areas | |||||||||||||||||||||||||
Elevation | 3500-4000 m (11500-13100 ft) | |||||||||||||||||||||||||
Distribution |
AZ; CA; ID; NV; OR; UT; nw Mexico
|
CO
|
||||||||||||||||||||||||
Discussion | Subspecies 7 (7 in the flora). The Physaria kingii complex is in need of further study. It is widespread in the western United States, mostly in montane environments. This treatment recognizes a highly variable species with generally well-marked, geographically coherent subspecies. Hybridization may be involved in some of the subspecies, especially in subsp. kaibabensis, where molecular data indicate intra-individual genetic variation (pers. obs.). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||||||||||||||||||||||
Key |
|
|||||||||||||||||||||||||
Source | FNA vol. 7, p. 645. | FNA vol. 7, p. 624. | ||||||||||||||||||||||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||
Synonyms | Vesicaria kingii, Lesquerella kingii | |||||||||||||||||||||||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | Rollins: Brittonia 33: 339. (1981) | ||||||||||||||||||||||||
Web links |