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mid-bladderpod

denseflower bladderpod, low bladderpod

Habit Perennials; caudex (buried), branched, (thickened with persistent leaf bases, cespitose); densely pubescent (usually grayish-green), trichomes (sessile or short stalked, spreading), several-rayed, rays furcate or bifurcate, slightly fused at base, (tuberculate or finely tuberculate). Annuals or biennials; caudex simple or branched, (relatively small, cespitose); densely pubescent, trichomes (spreading, sessile or short-stalked), 5–7-rayed, rays distinct and simple, (tuberculate, finely tubercled with a U-shaped notch on one side).
Stems

several from base, erect to decumbent, (unbranched, stout, densely leafy sterile shoots sometimes present), (0.5–)4–2.5 dm.

simple or few to several from base, erect or decumbent, (rarely branched, usually leafy), to 4 dm.

Basal leaves

(clustered at stem base);

blade linear to linear-oblanceolate, 2–5 cm, margins entire, usually involute, sometimes flattened, (apex obtuse to subacute).

blade lyrate-pinnatifid, 1–7 cm, margins entire or shallowly dentate.

Cauline leaves

blade linear-oblanceolate to linear, 1–3.5(–4.5) cm, margins entire, usually involute.

(sessile or shortly petiolate);

blade narrowly obovate to elliptic, 0.5–6 cm, margins entire, repand, or shallowly dentate.

Racemes

compact, (often nearly subumbellate).

dense, (elongated in fruit, often subtended by distal cauline leaves).

Flowers

sepals (yellowish or greenish yellow), ovate or oblong, 4.5–7.5(–9) mm, (lateral pair sometimes cucullate, median pair tapering at both ends, thickened apically, cucullate);

petals spatulate or oblong, 6.5–10.5(–15) mm, (base sometimes widened, apex rounded or retuse).

sepals elliptic, 3.7–7.2 mm, (lateral pair somewhat cucullate, median pair thickened apically);

petals (yellow to orange-yellow), obovate to obdeltate, (4.5–)7–10(–11) mm, (tapering to short claw, apex often emarginate).

Fruiting pedicels

(often expanded distally, ascending or recurved, usually straight or slightly curved, rarely nearly sigmoid), 4–15 mm, (stout).

(usually divaricate-spreading, straight or slightly curved, delicate, sometimes drooping, especially on herbarium specimens), 7–10 mm, (somewhat rigid).

Fruits

(sessile or substipitate), subglobose to slightly ovoid, usually inflated, rarely compressed or obcompressed, 4–6(–10) mm, (apex acute, slightly flattened);

valves sparsely pubescent, trichomes appressed;

ovules (8–)12–16(–20) per ovary;

style (2–)3–4.5(–5.5) mm.

(sessile or substipitate), globose or broadly obovate, not inflated, 4–6 mm, (smooth);

valves (not retaining seeds after dehiscence), glabrous;

replum as wide as or wider than fruit;

ovules 8–16 per ovary;

style 2–5 mm.

Seeds

flattened.

flattened.

2n

= 18, 20, 36.

= 14.

Physaria intermedia

Physaria densiflora

Phenology Flowering Apr–Aug. Flowering Mar–May.
Habitat Dry sandy, gravelly, or rocky soil, claylike hillsides, open chiprock, dry stream beds, gravel bars, open knolls, open pinyon-juniper woods, open stands of sagebrush, Gambel oak or ponderosa pine communities, calcareous substrates Sandy, granitic, or calcareous soils, sandy ledges, limestone outcrops, rocky prairies, uplands
Elevation 1600-2400 m (5200-7900 ft) 30-400 m (100-1300 ft)
Distribution
from FNA
AZ; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
TX
[BONAP county map]
Discussion

N. H. Holmgren (2005b) pointed out that the lectotype and other material from New Mexico, where Physaria intermedia is very infrequent, is quite similar to P. parvula from northern Colorado and northeastern Utah; it is also quite similar to, but less robust than, P. pulvinata from southwestern Colorado. The material from Arizona, northwestern New Mexico, and Utah may represent an unnamed taxon; further study is needed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Alyssum densiflorum (A. Gray) Kuntze (1891), not Desfontaines (1808) is an illegitimate name, sometimes found in synonymy with Physaria densiflora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 644. FNA vol. 7, p. 633.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella alpina var. intermedia, Lesquerella intermedia Vesicaria densiflora, Lesquerella densiflora
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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