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mid-bladderpod

Rocky Mountain bladderpod

Habit Perennials; caudex (buried), branched, (thickened with persistent leaf bases, cespitose); densely pubescent (usually grayish-green), trichomes (sessile or short stalked, spreading), several-rayed, rays furcate or bifurcate, slightly fused at base, (tuberculate or finely tuberculate). Perennials; (compact); caudex branched; densely (silvery) pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays distinct, furcate or bifurcate, (umbonate, tuberculate and the center less so).
Stems

several from base, erect to decumbent, (unbranched, stout, densely leafy sterile shoots sometimes present), (0.5–)4–2.5 dm.

several from base, erect or outer ones decumbent, (unbranched, stout, usually sparsely leaved), 1–3 dm.

Basal leaves

(clustered at stem base);

blade linear to linear-oblanceolate, 2–5 cm, margins entire, usually involute, sometimes flattened, (apex obtuse to subacute).

blade linear, 2–7(–10) cm, margins entire, repand, or shallowly dentate.

Cauline leaves

blade linear-oblanceolate to linear, 1–3.5(–4.5) cm, margins entire, usually involute.

(sessile);

blade (erect), spatulate to linear, (1–)2–3(–4.5) cm, margins entire, sometimes involute, (apex acute or subacute).

Racemes

compact, (often nearly subumbellate).

dense, (exceeding basal leaves).

Flowers

sepals (yellowish or greenish yellow), ovate or oblong, 4.5–7.5(–9) mm, (lateral pair sometimes cucullate, median pair tapering at both ends, thickened apically, cucullate);

petals spatulate or oblong, 6.5–10.5(–15) mm, (base sometimes widened, apex rounded or retuse).

sepals ovate or oblong, (4.5–)5–6(–7) mm, (lateral pair subsaccate, cucullate, median pair thickened, cucullate apically);

petals spatulate, 7–9(–11) mm (widened at base, slightly retuse).

Fruiting pedicels

(often expanded distally, ascending or recurved, usually straight or slightly curved, rarely nearly sigmoid), 4–15 mm, (stout).

(spreading, sharply sigmoid), 8–15 mm.

Fruits

(sessile or substipitate), subglobose to slightly ovoid, usually inflated, rarely compressed or obcompressed, 4–6(–10) mm, (apex acute, slightly flattened);

valves sparsely pubescent, trichomes appressed;

ovules (8–)12–16(–20) per ovary;

style (2–)3–4.5(–5.5) mm.

(sessile or substipitate), ovate to oblong, not compressed at distal margins or apex, 5–9 mm;

valves sparsely pubescent, trichomes appressed;

ovules 4–8 per ovary;

style 3–5 mm.

Seeds

flattened.

flattened.

2n

= 18, 20, 36.

= 16, ca. 20.

Physaria intermedia

Physaria calcicola

Phenology Flowering Apr–Aug. Flowering May–Jun.
Habitat Dry sandy, gravelly, or rocky soil, claylike hillsides, open chiprock, dry stream beds, gravel bars, open knolls, open pinyon-juniper woods, open stands of sagebrush, Gambel oak or ponderosa pine communities, calcareous substrates Shale bluffs, limestone hillsides, gypseous knolls and ravines, calcareous substrates, grasslands and pinyon-juniper communities
Elevation 1600-2400 m (5200-7900 ft) 1400-2100 m (4600-6900 ft)
Distribution
from FNA
AZ; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; NM
[WildflowerSearch map]
[BONAP county map]
Discussion

N. H. Holmgren (2005b) pointed out that the lectotype and other material from New Mexico, where Physaria intermedia is very infrequent, is quite similar to P. parvula from northern Colorado and northeastern Utah; it is also quite similar to, but less robust than, P. pulvinata from southwestern Colorado. The material from Arizona, northwestern New Mexico, and Utah may represent an unnamed taxon; further study is needed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 644. FNA vol. 7, p. 629.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella alpina var. intermedia, Lesquerella intermedia Lesquerella calcicola
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) (Rollins) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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