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mid-bladderpod

golden bladderpod

Habit Perennials; caudex (buried), branched, (thickened with persistent leaf bases, cespitose); densely pubescent (usually grayish-green), trichomes (sessile or short stalked, spreading), several-rayed, rays furcate or bifurcate, slightly fused at base, (tuberculate or finely tuberculate). Biennials or perennials; (short-lived); caudex branched; densely pubescent, trichomes (sessile or short-stalked, simple or not), 5–9-rayed, rays furcate, (fine, smooth or finely tuberculate).
Stems

several from base, erect to decumbent, (unbranched, stout, densely leafy sterile shoots sometimes present), (0.5–)4–2.5 dm.

several from base, erect or outer ones decumbent or procumbent, (sometimes much-branched distally), to 6 dm.

Basal leaves

(clustered at stem base);

blade linear to linear-oblanceolate, 2–5 cm, margins entire, usually involute, sometimes flattened, (apex obtuse to subacute).

blade obovate or rhombic, to ca. 2.5 cm, margins usually shallowly dentate, sometimes lyrate-pinnatifid.

Cauline leaves

blade linear-oblanceolate to linear, 1–3.5(–4.5) cm, margins entire, usually involute.

(proximal shortly petiolate, distal sessile);

blade obovate to rhombic or oblanceolate, 2–4(–6) cm, margins entire or shallowly and remotely dentate.

Racemes

compact, (often nearly subumbellate).

usually dense, (several-flowered).

Flowers

sepals (yellowish or greenish yellow), ovate or oblong, 4.5–7.5(–9) mm, (lateral pair sometimes cucullate, median pair tapering at both ends, thickened apically, cucullate);

petals spatulate or oblong, 6.5–10.5(–15) mm, (base sometimes widened, apex rounded or retuse).

sepals ovate or oblong (tapering at base), 3.6–4.8(–5.3) mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals obovate to spatulate, 4.5–7.5 mm, (blade narrowed to broad claw, margins sinuate).

Fruiting pedicels

(often expanded distally, ascending or recurved, usually straight or slightly curved, rarely nearly sigmoid), 4–15 mm, (stout).

(strongly recurved), to 20 mm.

Fruits

(sessile or substipitate), subglobose to slightly ovoid, usually inflated, rarely compressed or obcompressed, 4–6(–10) mm, (apex acute, slightly flattened);

valves sparsely pubescent, trichomes appressed;

ovules (8–)12–16(–20) per ovary;

style (2–)3–4.5(–5.5) mm.

(± pendent), ovoid, obcompressed, or globose, compressed, 4–6(–8) mm;

valves (not retaining seeds after dehiscence), sparsely pubescent or glabrous, sparsely pubescent inside;

replum as wide as or wider than fruit;

ovules usually 4, rarely 6, per ovary;

style 2.5–3.6 mm.

Seeds

flattened.

flattened.

2n

= 18, 20, 36.

= 14.

Physaria intermedia

Physaria aurea

Phenology Flowering Apr–Aug. Flowering Jun–Aug.
Habitat Dry sandy, gravelly, or rocky soil, claylike hillsides, open chiprock, dry stream beds, gravel bars, open knolls, open pinyon-juniper woods, open stands of sagebrush, Gambel oak or ponderosa pine communities, calcareous substrates Open sites and bare areas in rocky limestone soil in mountains, roadbanks, open woods
Elevation 1600-2400 m (5200-7900 ft) 2000-2800 m (6600-9200 ft)
Distribution
from FNA
AZ; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
NM
[BONAP county map]
Discussion

N. H. Holmgren (2005b) pointed out that the lectotype and other material from New Mexico, where Physaria intermedia is very infrequent, is quite similar to P. parvula from northern Colorado and northeastern Utah; it is also quite similar to, but less robust than, P. pulvinata from southwestern Colorado. The material from Arizona, northwestern New Mexico, and Utah may represent an unnamed taxon; further study is needed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Physaria aurea (known from the Jicarilla and Sacramento mountains) is similar to 35. P. gooddingii, which is found farther west in the mountains of Catron and Sierra counties, New Mexico, and Greenlee County, Arizona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 644. FNA vol. 7, p. 628.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella alpina var. intermedia, Lesquerella intermedia Lesquerella aurea
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) (Wooton) O’Kane & Al-Shehbaz: Novon 12: 322. (2002)
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