Physaria intermedia |
Physaria arenosa |
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mid-bladderpod |
Great Plains bladderpod |
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Habit | Perennials; caudex (buried), branched, (thickened with persistent leaf bases, cespitose); densely pubescent (usually grayish-green), trichomes (sessile or short stalked, spreading), several-rayed, rays furcate or bifurcate, slightly fused at base, (tuberculate or finely tuberculate). | Perennials or, rarely, annuals; caudex simple or branched; ± densely pubescent, trichomes (sessile or short-stalked), few-rayed, rays (usually spreading), distinct or slightly fused at base, furcate or bifurcate, (tuberculate). | ||||
Stems | several from base, erect to decumbent, (unbranched, stout, densely leafy sterile shoots sometimes present), (0.5–)4–2.5 dm. |
simple or few from base, prostrate or straggling to erect, (sometimes purplish, usually unbranched), (0.5–)1–2(–3) dm. |
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Basal leaves | (clustered at stem base); blade linear to linear-oblanceolate, 2–5 cm, margins entire, usually involute, sometimes flattened, (apex obtuse to subacute). |
blade oblanceolate, 1.5–5(–7) cm, margins entire or shallowly dentate, (flat). |
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Cauline leaves | blade linear-oblanceolate to linear, 1–3.5(–4.5) cm, margins entire, usually involute. |
blade elliptic to linear, (0.5–)1–2.5(–3) cm, margins usually entire. |
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Racemes | compact, (often nearly subumbellate). |
(secund), dense, (elongated in fruit). |
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Flowers | sepals (yellowish or greenish yellow), ovate or oblong, 4.5–7.5(–9) mm, (lateral pair sometimes cucullate, median pair tapering at both ends, thickened apically, cucullate); petals spatulate or oblong, 6.5–10.5(–15) mm, (base sometimes widened, apex rounded or retuse). |
sepals elliptic or oblong, 4–6(–7) mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (often red or lavender when dried), obovate, 6–8.5(–9.5) mm, (narrowing to broad claw). |
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Fruiting pedicels | (often expanded distally, ascending or recurved, usually straight or slightly curved, rarely nearly sigmoid), 4–15 mm, (stout). |
(usually sharply recurved, sometimes divaricate-spreading or nearly horizontal), 5–15(–20) mm, (stout). |
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Fruits | (sessile or substipitate), subglobose to slightly ovoid, usually inflated, rarely compressed or obcompressed, 4–6(–10) mm, (apex acute, slightly flattened); valves sparsely pubescent, trichomes appressed; ovules (8–)12–16(–20) per ovary; style (2–)3–4.5(–5.5) mm. |
subglobose, obovoid, or broadly ellipsoid, slightly inflated, (3.5–)4–5.5(–6.5) mm; valves densely pubescent outside, trichomes spreading or closely appressed, rarely sparsely pubescent inside; ovules (4–)8(–10) per ovary; style (slender), 3–5.5(–6.5) mm. |
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Seeds | flattened. |
slightly flattened. |
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2n | = 18, 20, 36. |
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Physaria intermedia |
Physaria arenosa |
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Phenology | Flowering Apr–Aug. | |||||
Habitat | Dry sandy, gravelly, or rocky soil, claylike hillsides, open chiprock, dry stream beds, gravel bars, open knolls, open pinyon-juniper woods, open stands of sagebrush, Gambel oak or ponderosa pine communities, calcareous substrates | |||||
Elevation | 1600-2400 m (5200-7900 ft) | |||||
Distribution |
AZ; NM; UT
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CO; MT; ND; NE; SD; WY; AB; MB; SK
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Discussion | N. H. Holmgren (2005b) pointed out that the lectotype and other material from New Mexico, where Physaria intermedia is very infrequent, is quite similar to P. parvula from northern Colorado and northeastern Utah; it is also quite similar to, but less robust than, P. pulvinata from southwestern Colorado. The material from Arizona, northwestern New Mexico, and Utah may represent an unnamed taxon; further study is needed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 644. | FNA vol. 7, p. 626. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Lesquerella alpina var. intermedia, Lesquerella intermedia | Vesicaria arenosa, Lesquerella arenosa, Lesquerella argentea var. arenosa, Lesquerella ludoviciana var. arenosa | ||||
Name authority | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) | (Richardson) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) | ||||
Web links |