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mid-bladderpod

threadleaf bladderpod

Habit Perennials; caudex (buried), branched, (thickened with persistent leaf bases, cespitose); densely pubescent (usually grayish-green), trichomes (sessile or short stalked, spreading), several-rayed, rays furcate or bifurcate, slightly fused at base, (tuberculate or finely tuberculate). Annuals; with a fine taproot; ± densely pubescent, trichomes several-rayed, rays distinct or fused at base, bifurcate, (prominently tuberculate throughout).
Stems

several from base, erect to decumbent, (unbranched, stout, densely leafy sterile shoots sometimes present), (0.5–)4–2.5 dm.

simple or few to several from base, erect, (sometimes branched), to 4 dm.

Basal leaves

(clustered at stem base);

blade linear to linear-oblanceolate, 2–5 cm, margins entire, usually involute, sometimes flattened, (apex obtuse to subacute).

blade elliptic to rhombic, 3–8 cm, (base narrowing gradually to petiole), margins entire, repand, coarsely toothed, or pinnatifid.

Cauline leaves

blade linear-oblanceolate to linear, 1–3.5(–4.5) cm, margins entire, usually involute.

(proximal often shortly petiolate, distal sessile);

blade linear or narrowly obovate, 1.5–6(–10) cm, margins entire, repand, or shallowly toothed.

Racemes

compact, (often nearly subumbellate).

usually loose.

Flowers

sepals (yellowish or greenish yellow), ovate or oblong, 4.5–7.5(–9) mm, (lateral pair sometimes cucullate, median pair tapering at both ends, thickened apically, cucullate);

petals spatulate or oblong, 6.5–10.5(–15) mm, (base sometimes widened, apex rounded or retuse).

sepals elliptic or ovate, 4–6 mm, (lateral pair usually subsaccate);

petals obovate to obdeltate, 6–10 mm, (apex often emarginate).

Fruiting pedicels

(often expanded distally, ascending or recurved, usually straight or slightly curved, rarely nearly sigmoid), 4–15 mm, (stout).

(usually divaricate, sometimes horizontal, straight or slightly curved), 8–20 mm.

Fruits

(sessile or substipitate), subglobose to slightly ovoid, usually inflated, rarely compressed or obcompressed, 4–6(–10) mm, (apex acute, slightly flattened);

valves sparsely pubescent, trichomes appressed;

ovules (8–)12–16(–20) per ovary;

style (2–)3–4.5(–5.5) mm.

not didymous, ± globose, slightly inflated, 4–6 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4 per ovary;

style 2–3.5 mm; (stigma expanded).

Seeds

flattened.

flattened, (margined).

2n

= 18, 20, 36.

= 10.

Physaria intermedia

Physaria angustifolia

Phenology Flowering Apr–Aug. Flowering Apr(-May).
Habitat Dry sandy, gravelly, or rocky soil, claylike hillsides, open chiprock, dry stream beds, gravel bars, open knolls, open pinyon-juniper woods, open stands of sagebrush, Gambel oak or ponderosa pine communities, calcareous substrates Shallow limestone-derived soils, sometimes spreading to disturbed sites
Elevation 1600-2400 m (5200-7900 ft) 90-300 m (300-1000 ft)
Distribution
from FNA
AZ; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
OK; TX
[BONAP county map]
Discussion

N. H. Holmgren (2005b) pointed out that the lectotype and other material from New Mexico, where Physaria intermedia is very infrequent, is quite similar to P. parvula from northern Colorado and northeastern Utah; it is also quite similar to, but less robust than, P. pulvinata from southwestern Colorado. The material from Arizona, northwestern New Mexico, and Utah may represent an unnamed taxon; further study is needed.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 644. FNA vol. 7, p. 625.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms Lesquerella alpina var. intermedia, Lesquerella intermedia Vesicaria angustifolia, Lesquerella angustifolia, Lesquerella longifolia
Name authority (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 324. (2002) (Nuttall ex Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 321. (2002)
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