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Hitchcock's bladderpod

Habit Plants forming soft mats; caudex elongated, elastic (creeping). Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate).
Stems

few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm.

Basal leaves

petiole differentiated from blade (sometimes weakly);

blade oblanceolate to obovate.

(petiole and blade differentiated or not);

blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire.

Cauline leaves

similar to basal, smaller.

Racemes

dense.

Flowers

sepals narrowly lanceolate to lanceolate, 2.8–6 mm;

petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade).

Fruiting pedicels

(ascending, straight or slightly curved), 2–6 mm.

Fruits

1.7–3 mm wide.

(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute);

valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–8 per ovary;

style 1.7–6 mm.

Seeds

flattened.

Anthers

1–1.5 mm.

Physaria hitchcockii subsp. confluens

Physaria hitchcockii

Phenology Flowering Jun–Jul.
Habitat Gravelly or rocky limestone
Elevation 2300-3500 m (7500-11500 ft)
Distribution
from FNA
NV
[BONAP county map]
from FNA
NV; UT
[BONAP county map]
Discussion

Subspecies confluens is found at or above timberline in the Grant, Quinn Canyon, and Schell Creek ranges.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudices elongated and elastic; basal leaves: petiole not differentiated from blade, blades linear-oblanceolate; Aquarius, Markagunt, and Paunsaugunt plateaus, Utah (limited to the pink member of the limestone Wasatch (Claron) Formation).
subsp. rubicundula
1. Caudices elongated or not, elastic or not; basal leaves: petiole differentiated (sometimes weakly) from blade, blades oblanceolate to obovate; Nevada, Utah
→ 2
2. Plants forming tufts; caudices not elongated, not elastic; fruits 2.6-3.8 mm wide; Table Cliff Plateau, Utah (limited to the white member of the limestone Wasatch (Claron) Formation) and limestones of the Sheep Range and Spring Mountains, Nevada.
subsp. hitchcockii
2. Plants forming soft mats; caudices elongated, elastic (creeping); fruits 1.7-3 mm wide; Grant, Quinn Canyon, and Schell Creek ranges, Nevada.
subsp. confluens
Source FNA vol. 7, p. 643. FNA vol. 7, p. 643.
Parent taxa Brassicaceae > tribe Physarieae > Physaria > Physaria hitchcockii Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. hitchcockii subsp. hitchcockii, P. hitchcockii subsp. rubicundula
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. hitchcockii subsp. confluens, P. hitchcockii subsp. hitchcockii, P. hitchcockii subsp. rubicundula
Synonyms Lesquerella hitchcockii subsp. confluens, Lesquerella confluens Lesquerella hitchcockii
Name authority (Maguire & A. H. Holmgren) O’Kane: Novon 17: 380. (2007) (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002)
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