Physaria gracilis |
Physaria thamnophila |
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spreading bladderpod |
Zapata bladderpod |
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Habit | Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). | Perennials; caudex simple or branched, (woody); densely (silvery) pubescent, trichomes (short-stalked), 4–8-rayed, rays furcate or bifurcate, (finely tuberculate throughout). | ||||
Stems | simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm. |
simple or few from base, decumbent, (straggling and flexuous, usually branched distally), 4–8 dm. |
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Basal leaves | blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent). |
blade narrowly elliptic to oblanceolate, 4–12 cm, margins entire, sinuate, or shallowly dentate, (apex acute). |
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Cauline leaves | (proximal petiolate, distal sessile); blade oblanceolate to oblong, 1–7 cm, margins dentate to repand. |
(sessile or proximal shortly petiolate); blade linear to narrowly elliptic, 3–4 cm, margins entire, sinuate, or remotely dentate, (apex acute). |
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Racemes | loose, (elongated). |
loose, (sometimes greatly elongated). |
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Flowers | sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate); petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw). |
sepals elliptic, 3.5–4 mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals broadly obovate, 4–5 mm, (sometimes with short, broad claw). |
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Fruiting pedicels | (usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout). |
(recurved), 15–20(–25) mm. |
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Fruits | (stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside; replum as wide as or wider than fruit; ovules 8–20(–28) per ovary; style 2–4.5 mm. |
(pendent), subglobose or broadly ovoid, slightly compressed, 5–7 mm; valves (not retaining seeds after dehiscence), glabrous throughout; ovules per ovary unknown; style 1.5–2 mm. |
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Seeds | slightly flattened. |
flattened. |
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2n | = 16. |
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Physaria gracilis |
Physaria thamnophila |
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Phenology | Flowering Apr. | |||||
Habitat | Sandy soils, entangled in shrubs, cactus clumps | |||||
Elevation | 1700-1800 m (5600-5900 ft) | |||||
Distribution |
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
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TX |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria thamnophila is found in sandy areas with shrubs and cactus in sparse shrubland communities of Starr and Zapata counties. It sometimes flowers through September with sufficient moisture. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 641. | FNA vol. 7, p. 664. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis | Lesquerella thamnophila | ||||
Name authority | (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 328. (2002) | ||||
Web links |