Physaria gracilis |
Physaria obcordata |
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spreading bladderpod |
Dudley bluffs twinpod, Piceance twinpod |
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Habit | Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). | Perennials; caudex branched, (thick, covered with persistent, overlapping leaf bases); densely (silvery) pubescent, trichomes (stellate-scalelike), several-rayed, rays fused (webbed) to tips. | ||||
Stems | simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm. |
several from base, erect to decumbent, (unbranched, fertile stems from among basal leaves), 1.2–1.8 dm. |
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Basal leaves | blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent). |
(erect, not rosulate); blade broadly oblanceolate, 4–8 cm, margins entire or shallowly sinuate-dentate, (apex acute). |
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Cauline leaves | (proximal petiolate, distal sessile); blade oblanceolate to oblong, 1–7 cm, margins dentate to repand. |
(proximal petiolate); blade narrowly lanceolate, similar to basal in size, (distal with cuneate base), margins entire. |
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Racemes | loose, (elongated). |
loose, (elongated in fruit). |
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Flowers | sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate); petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw). |
sepals (greenish yellow), elliptic, often broadly so, 4.8–7.1 mm; petals oblanceolate, 6.8–9.8 mm. |
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Fruiting pedicels | (usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout). |
(widely spreading to recurved), 1–1.5 cm. |
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Fruits | (stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside; replum as wide as or wider than fruit; ovules 8–20(–28) per ovary; style 2–4.5 mm. |
(usually pendent), slightly didymous apically, obcordate, slightly inflated, 4–7 × 3–6 mm, (papery, basal sinus absent, apical sinus evident to nearly absent); valves (retaining seeds after dehiscence), pubescent; replum broadly obovate to suborbicular, 4–5 mm, as wide as or wider than fruit, margins entire; ovules usually 4 (rarely 6–8) per ovary; style (2.5–)3–4(–5) mm. |
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Seeds | slightly flattened. |
plump. |
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Physaria gracilis |
Physaria obcordata |
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Phenology | Flowering May–Jun. | |||||
Habitat | Steep slopes, fine chiprock, shaley hillsides | |||||
Elevation | 1800-2300 m (5900-7500 ft) | |||||
Distribution |
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
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CO
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria obcordata is known from the Thirteenmile Creek Tongue and the Parachute Creek Member of the Green River Formation. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 641. | FNA vol. 7, p. 653. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis | |||||
Name authority | (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | Rollins: J. Arnold Arbor. 64: 495, fig. 1. (1983) | ||||
Web links |