Physaria gracilis |
Physaria nelsonii |
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spreading bladderpod |
Nelson's bladderpod |
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Habit | Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). | Perennials; (diminutive); caudex branched, (densely cespitose, mound-forming); densely pubescent, trichomes (flaring, giving a shaggy appearance), usually 4–5-rayed, rays furcate or bifurcate, (strongly tuberculate throughout). | ||||
Stems | simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm. |
few to several from base, erect, (from basal tuft), 0.1–0.2(–0.3) dm, (not or just barely exceeding leaves). |
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Basal leaves | blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent). |
blade spatulate to oblanceolate, 0.5–1.5 cm, (base gradually narrowed to petiole), margins entire. |
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Cauline leaves | (proximal petiolate, distal sessile); blade oblanceolate to oblong, 1–7 cm, margins dentate to repand. |
(absent or few); similar to basal, blade linear. |
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Racemes | loose, (elongated). |
dense, (few-flowered). |
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Flowers | sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate); petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw). |
sepals (pale yellow), oblong to elliptic, 4–5 mm, (median pair usually thickened apically, cucullate); petals (bright yellow), lingulate, 6–8 mm. |
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Fruiting pedicels | (usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout). |
(loosely sigmoid), 3–5 mm. |
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Fruits | (stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside; replum as wide as or wider than fruit; ovules 8–20(–28) per ovary; style 2–4.5 mm. |
lanceolate, compressed apically, 2.5–3(–4) mm; valves pubescent, trichomes spreading, (appearing fuzzy), rarely with trichomes inside; ovules 4–8 per ovary; style 2.5–4 mm. |
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Seeds | slightly flattened. |
plump, (oblong). |
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Physaria gracilis |
Physaria nelsonii |
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Phenology | Flowering May-mid Jun. | |||||
Habitat | Limestone, windswept knolls and cliffs, nearly barren areas with other cushion-forming plants | |||||
Elevation | 1600-2300 m (5200-7500 ft) | |||||
Distribution |
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
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UT; WY |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria nelsonii is morphologically similar to 73. P. pycnantha, which traditionally was included in a broader P. nelsonii. These allopatric species are distinguished by styles equal to or exceeding the length of fruit and fruiting stems overtopped by basal leaves (P. nelsonii) versus styles shorter than fruits and fruiting stems usually exserted beyond basal leaves (P. pycnantha). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 641. | FNA vol. 7, p. 652. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis | Lesquerella condensata, Lesquerella alpina subsp. condensata, Lesquerella alpina var. condensata | ||||
Name authority | (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | ||||
Web links |