Physaria gracilis |
Physaria macrocarpa |
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spreading bladderpod |
largefruit bladderpod |
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Habit | Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). | Perennials; caudex branched; densely pubescent, trichomes (sessile or short-stalked), 4–6-rayed, rays distinct, usually furcate, rarely bifurcate, (finely tuberculate throughout). | ||||
Stems | simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm. |
few or several from base, prostrate to decumbent, (unbranched or branched), 0.5–1.5 dm. |
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Basal leaves | blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent). |
blades orbicular to broadly obovate, 1.5–3 cm, margins usually entire, rarely remotely dentate. |
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Cauline leaves | (proximal petiolate, distal sessile); blade oblanceolate to oblong, 1–7 cm, margins dentate to repand. |
(sessile or shortly petiolate); blade elliptic to oblanceolate, 1–1.5(–2.5) cm, margins entire, (apex obtuse). |
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Racemes | loose, (elongated). |
dense, (elongated in fruit). |
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Flowers | sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate); petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw). |
sepals ovate or oblong-elliptic, 5–5.5 mm, (lateral pair not saccate); petals cuneate or broadly obovate, ca. 7 mm, (sometimes slightly narrowed to a broad claw, apex sometimes retuse). |
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Fruiting pedicels | (usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout). |
(sharply recurved), 5–10 mm, (stout). |
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Fruits | (stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside; replum as wide as or wider than fruit; ovules 8–20(–28) per ovary; style 2–4.5 mm. |
subglobose to broadly obovoid, strongly inflated (often slightly angustiseptate), 5–7 mm, (papery); valves sparsely pubescent; (septum fenestrate, perforate, or obsolete); ovules 4–8 per ovary; style 2–3 mm. |
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Seeds | slightly flattened. |
somewhat flattened. |
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Physaria gracilis |
Physaria macrocarpa |
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Phenology | Flowering May–Jun. | |||||
Habitat | Gypsum-clay hills and benches, naked clay flats and barren hills | |||||
Elevation | 2000-2400 m (6600-7900 ft) | |||||
Distribution |
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
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WY |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria macrocarpa is found in the Great Divide and Green River basins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 641. | FNA vol. 7, p. 650. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis | Lesquerella macrocarpa | ||||
Name authority | (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) | ||||
Web links |