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spreading bladderpod

Graham's twinpod

Habit Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate.
Stems

simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm.

several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm.

Basal leaves

blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent).

(outer ones spreading, inner erect or ascending);

blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed.

Cauline leaves

(proximal petiolate, distal sessile);

blade oblanceolate to oblong, 1–7 cm, margins dentate to repand.

similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous).

Racemes

loose, (elongated).

loose, (elongated).

Flowers

sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate);

petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw).

sepals lanceolate or narrowly oblong, 5.8–7.2 mm;

petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed).

Fruiting pedicels

(usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout).

(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm.

Fruits

(stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm;

valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside;

replum as wide as or wider than fruit;

ovules 8–20(–28) per ovary;

style 2–4.5 mm.

didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep);

valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy;

replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse;

ovules 4 per ovary;

style (4–)5–7 mm.

Seeds

slightly flattened.

plump, (suborbicular).

Physaria gracilis

Physaria grahamii

Phenology Flowering May–Jun.
Habitat Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay
Elevation 2100-2900 m (6900-9500 ft)
Distribution
from FNA
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems to 7 dm; cauline leaves: blade margins usually deeply dentate, rarely repand; fruits ± sessile, globose or subglobose, 3-6 mm, bases rounded.
subsp. gracilis
1. Stems usually less than 3 dm; cauline leaves: blade margins frequently repand, occasionally dentate; fruits stipitate (gynophore slender), obpyriform to narrowly obovoid, (5-)6-9 mm, bases truncate.
subsp. nuttallii
Source FNA vol. 7, p. 641. FNA vol. 7, p. 642.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. gracilis subsp. gracilis, P. gracilis subsp. nuttallii
Synonyms Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda
Name authority (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937)
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