Physaria gracilis |
Physaria grahamii |
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spreading bladderpod |
Graham's twinpod |
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Habit | Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). | Perennials; caudex branched, (thick, cespitose); densely pubescent, trichomes rays (appressed on leaves, ascending on pedicels and fruits), distinct, furcate or bifurcate. | ||||
Stems | simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm. |
several from base, decumbent to erect or ascending (unbranched), 1–2.5 dm. |
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Basal leaves | blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent). |
(outer ones spreading, inner erect or ascending); blade ovate, often broadly so, 4–7 cm, margins repand to lyrate-lobed. |
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Cauline leaves | (proximal petiolate, distal sessile); blade oblanceolate to oblong, 1–7 cm, margins dentate to repand. |
similar to basal, blade oblanceolate or narrowly oblong, reduced in size, (base gibbous). |
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Racemes | loose, (elongated). |
loose, (elongated). |
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Flowers | sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate); petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw). |
sepals lanceolate or narrowly oblong, 5.8–7.2 mm; petals (erect, sometime purplish or drying purple), narrowly oblong to oblanceolate, 7–10 mm, (not or weakly clawed). |
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Fruiting pedicels | (usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout). |
(ascending to divaricate-ascending, sigmoid to nearly straight), 10–17 mm. |
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Fruits | (stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside; replum as wide as or wider than fruit; ovules 8–20(–28) per ovary; style 2–4.5 mm. |
didymous, globose or subglobose, inflated, 10–13 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes ascending, appearing fuzzy; replum oblong to oblanceolate, as wide as or wider than fruit, apex obtuse; ovules 4 per ovary; style (4–)5–7 mm. |
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Seeds | slightly flattened. |
plump, (suborbicular). |
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Physaria gracilis |
Physaria grahamii |
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Phenology | Flowering May–Jun. | |||||
Habitat | Sagebrush, pinyon-juniper, ponderosa pine, Douglas-fir, limber pine communities on clay, or a mixture of shale fragments and clay | |||||
Elevation | 2100-2900 m (6900-9500 ft) | |||||
Distribution |
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
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UT |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria grahamii is difficult to evaluate due to the paucity of collections. The tentative recognition by N. H. Holmgren (2005b) is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 641. | FNA vol. 7, p. 642. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis | P. acutifolia var. purpurea, P. acutifolia var. repanda, P. repanda | ||||
Name authority | (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | C. V. Morton: Ann. Carnegie Mus. 26: 220. (1937) | ||||
Web links |