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spreading bladderpod

Photo is of parent taxon

Nuttall's bladderpod

Habit Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate).
Stems

simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm.

usually less than 3 dm;

bud clusters erect.

Basal leaves

blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent).

Cauline leaves

(proximal petiolate, distal sessile);

blade oblanceolate to oblong, 1–7 cm, margins dentate to repand.

blade margins frequently repand, occasionally dentate.

Racemes

loose, (elongated).

Flowers

sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate);

petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw).

Fruiting pedicels

(usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout).

slender.

Fruits

(stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm;

valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside;

replum as wide as or wider than fruit;

ovules 8–20(–28) per ovary;

style 2–4.5 mm.

stipitate (gynophore slender), obpyriform to narrowly obovoid, (5–)6–9 mm, base truncate.

Seeds

slightly flattened.

2n

= 12.

Physaria gracilis

Physaria gracilis subsp. nuttallii

Phenology Flowering Apr–May.
Habitat Prairies, pastures, hillsides, rocky limestone, shallow soil of glades, roadsides, open fields
Elevation 150-500 m (500-1600 ft)
Distribution
from FNA
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
KS; OK; TX
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems to 7 dm; cauline leaves: blade margins usually deeply dentate, rarely repand; fruits ± sessile, globose or subglobose, 3-6 mm, bases rounded.
subsp. gracilis
1. Stems usually less than 3 dm; cauline leaves: blade margins frequently repand, occasionally dentate; fruits stipitate (gynophore slender), obpyriform to narrowly obovoid, (5-)6-9 mm, bases truncate.
subsp. nuttallii
Source FNA vol. 7, p. 641. FNA vol. 7, p. 641.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria > Physaria gracilis
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. gracilis subsp. gracilis
Subordinate taxa
P. gracilis subsp. gracilis, P. gracilis subsp. nuttallii
Synonyms Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis Vesicaria nuttallii, Alyssum nuttallii, Alyssum repandum, Lesquerella gracilis subsp. nuttallii, Lesquerella gracilis var. repanda, Lesquerella nuttallii, Lesquerella repanda, P. gracilis var. repanda, Vesicaria repanda
Name authority (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) (Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002)
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