Physaria gracilis |
Physaria globosa |
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spreading bladderpod |
globe bladderpod |
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Habit | Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). | Biennials or perennials; caudex branched, (± woody); densely pubescent, trichomes (sessile), 3–6-rayed, rays distinct and simple or furcate, (in 2 layers, lower layer umbonate, smooth to finely tuberculate, some often with a U-shaped notch). | ||||
Stems | simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm. |
several from base, erect, (arising among leaves of an elongated main axis), to 5 dm (± equal). |
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Basal leaves | blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent). |
(shortly petiolate); blade obovate to oblanceolate, (1.5–)2.5–5(–6) cm, margins entire, sinuate to shallowly toothed, or pinnatifid. |
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Cauline leaves | (proximal petiolate, distal sessile); blade oblanceolate to oblong, 1–7 cm, margins dentate to repand. |
(sessile or shortly petiolate); blade oblanceolate to oblong, 1.3–3(–4) cm, (base cuneate), margins entire or repand to dentate. |
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Racemes | loose, (elongated). |
dense. |
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Flowers | sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate); petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw). |
sepals elliptic or obovate, 2.6–4.1 mm, (median pair thickened apically, cucullate); petals (bright yellow), obovate, 3.5–6.5(–7.5) mm, (margins sinuate). |
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Fruiting pedicels | (usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout). |
(usually spreading horizontally, straight), 7–14(–21) mm. |
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Fruits | (stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside; replum as wide as or wider than fruit; ovules 8–20(–28) per ovary; style 2–4.5 mm. |
(sessile or substipitate); globose, often slightly compressed apically, (1–)2–3 mm; valves sparsely pubescent, sometimes pubescent inside, trichomes spreading, 3–5-rayed; ovules 4 per ovary; style 2–3.5(–4) mm. |
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Seeds | slightly flattened. |
flattened or plump, (often outer surface hemispherical, inner surface flattened, or both surfaces rounded). |
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2n | = 14. |
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Physaria gracilis |
Physaria globosa |
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Phenology | Flowering Mar–May. | |||||
Habitat | Open rocky areas, shale at cliff bases, open talus, ledges, open cedar glades | |||||
Elevation | 100-300 m (300-1000 ft) | |||||
Distribution |
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
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IN; KY; TN |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria globosa is possibly introduced in Indiana. A report for Ohio was based on a collection by “Jones,” but that specimen cannot be located. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 641. | FNA vol. 7, p. 639. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis | Vesicaria globosa, Alyssum globosum, Alyssum shortii, Lesquerella globosa | ||||
Name authority | (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | (Desvaux) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | ||||
Web links |