Physaria gracilis |
Physaria eburniflora |
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spreading bladderpod |
Devils Gate twinpod |
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Habit | Annuals or biennials; (delicate, wiry); with a fine taproot; pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays distinct, usually furcate, occasionally bifurcate, (smooth to somewhat tuberculate). | Perennials; caudex usually simple; densely pubescent, trichomes (sessile), rays often furcate, fused toward base, (nearly smooth). | ||||
Stems | simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm. |
simple from base, prostrate, (arising lateral to rosette), 0.1–0.5 dm. |
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Basal leaves | blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent). |
blade suborbicular, (1–)2.5(–3) cm, (base abruptly narrowed to petiole), margins entire, (flat), (surfaces densely silvery pubescent, trichomes in multiple layers, appressed). |
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Cauline leaves | (proximal petiolate, distal sessile); blade oblanceolate to oblong, 1–7 cm, margins dentate to repand. |
(2–4); blade oblanceolate, ca. 1 cm, (base cuneate), margins entire, (apex acute). |
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Racemes | loose, (elongated). |
condensed. |
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Flowers | sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate); petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw). |
sepals (erect, purplish to greenish), linear-oblong or boat-shaped, 5.5–6.5 mm, (lateral pair more saccate than median); petals (white), spatulate, 9–12 mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout). |
(divaricate-ascending, nearly straight), 6–10 mm. |
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Fruits | (stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside; replum as wide as or wider than fruit; ovules 8–20(–28) per ovary; style 2–4.5 mm. |
strongly didymous, irregular in shape and size, (base slightly cordate, apex with a deep closed sinus), strongly to somewhat inflated, 6–8 × 6–8 mm (± bladderlike, papery); valves (retaining seeds after dehiscence), pubescent; replum elliptic to obovate, not constricted, as wide as or wider than fruit, apex obtuse; ovules 4–8 per ovary; style 4–5 mm, (sparsely pubescent or glabrous). |
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Seeds | slightly flattened. |
plump. |
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Physaria gracilis |
Physaria eburniflora |
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Phenology | Flowering May–Jun. | |||||
Habitat | Limestone hills, red soil, rocky calcareous slopes, clay depressions, granite and marble detritus | |||||
Elevation | 1800-3000 m (5900-9800 ft) | |||||
Distribution |
AL; IA; IL; KS; LA; MO; MS; OK; TN; TX
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WY |
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Discussion | Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 641. | FNA vol. 7, p. 635. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria gracilis, Alyssum gracile, Lesquerella gracilis | |||||
Name authority | (Hooker) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | Rollins: Brittonia 33: 333. (1981) | ||||
Web links |