Physaria gracilis
Physaria chambersii
spreading bladderpod
Chambers' bladder-pod, Chambers' physaria, Chambers' twinpod, double bladderpod
simple to several from base, erect, often outer decumbent, (unbranched or branched distally), 1–7 dm.
several from base, erect or decumbent (arising laterally, unbranched), 0.5–1.5 mm.
blade oblanceolate to elliptic, 1.5–8(–11.5) cm, margins lyrate-pinnatifid to dentate or repand, (abaxial surface densely pubescent, adaxial sparsely pubescent).
(petiole slender);
blade obovate to orbicular, 3–6 cm (width 10–20 mm), margins entire or dentate.
(proximal petiolate, distal sessile);
blade oblanceolate to oblong, 1–7 cm, margins dentate to repand.
blade spatulate, 1–2 cm (width 3–6 mm), margins entire, (apex often acute).
loose, (elongated).
congested.
sepals elliptic or broadly ovate, 3–6.5(–8) mm, (median pair slightly thickened apically, cucullate);
petals (yellow to orange), broadly obovate, 6–11 mm, (narrowing gradually to short claw).
sepals narrowly lanceolate, 5–8(–9) mm;
petals narrowly oblanceolate, 9–12 mm, (claw undifferentiated from blade).
(usually divaricate-spreading, sometimes horizontal or shallowly recurved, straight or slightly curved), (7–)10–20(–25) mm, (slender or stout).
(divaricate, slightly sigmoid), 8–15 mm.
(stipitate or subsessile, gynophore 1–2 mm), globose, subglobose, obpyriform, or obovoid, not or slightly inflated, 3–9 mm;
valves (not retaining seeds after dehiscence), glabrous throughout or sparsely pubescent inside;
replum as wide as or wider than fruit;
ovules 8–20(–28) per ovary;
style 2–4.5 mm.
(often purplish in age), didymous, subreniform, strongly inflated, 9–18 × 11–21(–30) mm, (papery, base obtuse to slightly cordate, apical sinus V-shaped or convex, open crests rounded);
valves (2-keeled on side away from replum, each 3-sided, keels rounded, sides flat or slightly convex, retaining seeds after dehiscence), evenly and densely pubescent;
replum oblong, as wide as or wider than fruits, apex obtuse;
ovules 4–12 per ovary;
style (4–)6–8 mm (exceeding sinus).
slightly flattened.
flattened.
= 8, 10, 16, 24.
Physaria gracilis
Physaria chambersii
Subspecies 2 (2 in the flora).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Physaria chambersii has been divided into three varieties based on whether the fruit is stipitate (var. canaani) or not, and whether the caudex elongates (var. sobolifera) or not (var. chambersii). In this species and in some others, e.g., P. newberryi, the latter character often depends on substrate and microclimate. Shifting substrates, such as moving sand and talus, often cause caudices to elongate. The species can be confused with 57. P. newberryi.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Stems to 7 dm; cauline leaves: blade margins usually deeply dentate, rarely repand; fruits ± sessile, globose or subglobose, 3-6 mm, bases rounded. | subsp. gracilis |
1. Stems usually less than 3 dm; cauline leaves: blade margins frequently repand, occasionally dentate; fruits stipitate (gynophore slender), obpyriform to narrowly obovoid, (5-)6-9 mm, bases truncate. | subsp. nuttallii |
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