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Gordon's bladderpod

Kane County twinpod

Habit Annuals, biennials, or perennials; (short-lived); with a fine taproot; usually densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays distinct and furcate or bifurcate, (nearly smooth to finely tuberculate). Perennials; caudex simple, (with deep roots, thickened); densely (silvery) pubescent throughout (densely covering leaves with several appressed layers), less dense on stems, trichomes (stellate-scalelike), rays fused (webbed) in proximal 1/2 or to tips, (umbonate, nearly smooth to moderately tuberculate).
Stems

several from base, erect to decumbent or prostrate, (unbranched or branched, sometimes densely leaved), 1–3.5(–4.5) dm.

simple from base, erect or outer ones slightly decumbent toward base, (from below or in basal leaves, unbranched), (0.5–)0.8–1.6(–2) dm.

Basal leaves

blade obovate to broadly oblong, 1.5–5(–8) cm, margins lyrate-pinnatifid, dentate, or entire.

(erect, petiole long, slender);

blade spatulate to broadly oblanceolate, (3–)5–7(–12) cm, (base gradually tapering to petiole), margins entire, (apex rounded or obtuse).

Cauline leaves

(proximal sometimes petiolate, distal sessile);

blade linear to oblanceolate, often falcate, 1–4(–7) cm, (proximal with base sometimes cuneate), margins entire, repand, or shallowly dentate.

blade oblanceolate, similar to basal, (base cuneate), margins entire.

Racemes

dense.

dense.

Flowers

sepals elliptic or oblong, 3–6.5 mm, (lateral pair subsaccate, median pair thickened apically, cucullate);

petals (widely spreading at anthesis, yellow to orange, claw sometimes whitish), cuneate, obdeltate, or obovate, (tapering to claw), 5–8(–10) mm, (claw often widened at base).

sepals (erect), linear to linear-oblong, somewhat boat-shaped, 7–10 mm;

petals (erect at anthesis), lingulate, 11–15 mm, (claw undifferentiated from blade).

Fruiting pedicels

(divaricate-ascending, sigmoid or, sometimes, nearly straight), 5–15(–25) mm.

(divaricate-ascending, straight or slightly curved), 10–15 mm.

Fruits

(shortly stipitate), subglobose, not or slightly compressed, (3–)4–8 mm;

valves (not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules (8–)12–20(–26) per ovary;

style (1.5–)2–4(–5) mm.

(purplish in age), strongly didymous, semiorbicular, highly inflated, 10–18 × 14–19 mm, (papery), basal sinus usually shallow, rarely absent, apical sinus deep, narrowly V-shaped;

valves retaining seeds after dehiscence, sides flat, back rounded, margins keeled, base and apex obtuse;

replum narrowly oblong to linear, as wide as or wider than fruit, base slightly narrowed, apex obtusely rounded;

ovules 4 per ovary;

style 3–5 mm, (slender).

Seeds

flattened.

slightly flattened.

2n

= 12, 32.

Physaria gordonii

Physaria lepidota

Phenology Flowering Feb–Jul.
Habitat Sandy or light soils, rocky plains, caprock ledges, gravelly brushland, sandy desert washes, stream bottoms, pastures, roadsides, abandoned fields
Elevation 150-1700 m (500-5600 ft)
Distribution
from FNA
AZ; KS; NM; OK; TX; VA; Mexico (Chihuahua, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Physaria gordonii was reported from Virginia in 1987 by Robert Wright from a Hampton Shale roadcut along the Blue Ridge Parkway, where it was probably a short-lived waif.

Subspecies densifolia, of Lincoln County, New Mexico, of which there is now more material than Rollins had available in 1993, appears to represent a suite of environmentally determined, variable, and intergrading characteristics that does not merit taxonomic recognition.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Trichomes: rays fused nearly to tips; fruits with deep sinuses, or shallow basally, deep apically.
subsp. lepidota
1. Trichomes: rays fused in proximal 1/2; fruits with sinuses absent or shallow basally, deep apically.
subsp. membranacea
Source FNA vol. 7, p. 640. FNA vol. 7, p. 648.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. lepidota subsp. lepidota, P. lepidota subsp. membranacea
Synonyms Vesicaria gordonii, Alyssum gordonii, Lesquerella gordonii, Lesquerella gordonii var. densifolia, P. gordonii subsp. densifolia, P. gordonii var. densifolia
Name authority (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) Rollins: Brittonia 33: 335, figs. 1, 2. (1981)
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