Physaria gordonii |
Physaria lepidota |
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Gordon's bladderpod |
Kane County twinpod |
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Habit | Annuals, biennials, or perennials; (short-lived); with a fine taproot; usually densely pubescent, trichomes (sessile or short-stalked), 4–7-rayed, rays distinct and furcate or bifurcate, (nearly smooth to finely tuberculate). | Perennials; caudex simple, (with deep roots, thickened); densely (silvery) pubescent throughout (densely covering leaves with several appressed layers), less dense on stems, trichomes (stellate-scalelike), rays fused (webbed) in proximal 1/2 or to tips, (umbonate, nearly smooth to moderately tuberculate). | ||||
Stems | several from base, erect to decumbent or prostrate, (unbranched or branched, sometimes densely leaved), 1–3.5(–4.5) dm. |
simple from base, erect or outer ones slightly decumbent toward base, (from below or in basal leaves, unbranched), (0.5–)0.8–1.6(–2) dm. |
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Basal leaves | blade obovate to broadly oblong, 1.5–5(–8) cm, margins lyrate-pinnatifid, dentate, or entire. |
(erect, petiole long, slender); blade spatulate to broadly oblanceolate, (3–)5–7(–12) cm, (base gradually tapering to petiole), margins entire, (apex rounded or obtuse). |
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Cauline leaves | (proximal sometimes petiolate, distal sessile); blade linear to oblanceolate, often falcate, 1–4(–7) cm, (proximal with base sometimes cuneate), margins entire, repand, or shallowly dentate. |
blade oblanceolate, similar to basal, (base cuneate), margins entire. |
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Racemes | dense. |
dense. |
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Flowers | sepals elliptic or oblong, 3–6.5 mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals (widely spreading at anthesis, yellow to orange, claw sometimes whitish), cuneate, obdeltate, or obovate, (tapering to claw), 5–8(–10) mm, (claw often widened at base). |
sepals (erect), linear to linear-oblong, somewhat boat-shaped, 7–10 mm; petals (erect at anthesis), lingulate, 11–15 mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (divaricate-ascending, sigmoid or, sometimes, nearly straight), 5–15(–25) mm. |
(divaricate-ascending, straight or slightly curved), 10–15 mm. |
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Fruits | (shortly stipitate), subglobose, not or slightly compressed, (3–)4–8 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules (8–)12–20(–26) per ovary; style (1.5–)2–4(–5) mm. |
(purplish in age), strongly didymous, semiorbicular, highly inflated, 10–18 × 14–19 mm, (papery), basal sinus usually shallow, rarely absent, apical sinus deep, narrowly V-shaped; valves retaining seeds after dehiscence, sides flat, back rounded, margins keeled, base and apex obtuse; replum narrowly oblong to linear, as wide as or wider than fruit, base slightly narrowed, apex obtusely rounded; ovules 4 per ovary; style 3–5 mm, (slender). |
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Seeds | flattened. |
slightly flattened. |
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2n | = 12, 32. |
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Physaria gordonii |
Physaria lepidota |
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Phenology | Flowering Feb–Jul. | |||||
Habitat | Sandy or light soils, rocky plains, caprock ledges, gravelly brushland, sandy desert washes, stream bottoms, pastures, roadsides, abandoned fields | |||||
Elevation | 150-1700 m (500-5600 ft) | |||||
Distribution |
AZ; KS; NM; OK; TX; VA; Mexico (Chihuahua, Sonora)
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UT |
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Discussion | Physaria gordonii was reported from Virginia in 1987 by Robert Wright from a Hampton Shale roadcut along the Blue Ridge Parkway, where it was probably a short-lived waif. Subspecies densifolia, of Lincoln County, New Mexico, of which there is now more material than Rollins had available in 1993, appears to represent a suite of environmentally determined, variable, and intergrading characteristics that does not merit taxonomic recognition. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 640. | FNA vol. 7, p. 648. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Vesicaria gordonii, Alyssum gordonii, Lesquerella gordonii, Lesquerella gordonii var. densifolia, P. gordonii subsp. densifolia, P. gordonii var. densifolia | |||||
Name authority | (A. Gray) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) | Rollins: Brittonia 33: 335, figs. 1, 2. (1981) | ||||
Web links |